Palaeontologia africana
Permanent URI for this communityhttps://wiredspace.wits.ac.za/handle/10539/13253
ISSN (print): 0078-8554
ISSN (electronic): 2410-4418
For queries regarding content of Palaeontologia africana collections please contact Jonah Choiniere by email : jonah.choiniere@wits.ac.za or Tel : 011 717 6684
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Item A new dicynodont (Anomodontia: Emydopoidea) from the terminal Permian of KwaZulu-Natal, South Africa(Evolutionary Studies Institute, 2019-04) Kammerer, Christian F.A new taxon of dicynodont (Thliptosaurus imperforatus gen. et sp. nov.) is described based on a dorsoventrally-crushed skull from latest Permian (upper Daptocephalus Assemblage Zone) strata in KwaZulu-Natal, South Africa. Thliptosaurus is distinguished from all other dicynodonts by an elongate intertemporal bar with broad dorsal exposure of the parietals but apparently no pineal foramen. Absence of the pineal foramen in dicynodonts is exceedingly rare; the only other taxa which exhibit this feature either have substantially broader (Kawingasaurus fossilis) or narrower (Kombuisia frerensis) intertemporal regions. Inclusion of Thliptosaurus in a phylogenetic analysis of dicynodonts recovers it as a kingoriid emydopoid, a position supported by its anteriorly-restricted pterygoid keel, elongate, curved anterior process of the lacrimal, relatively posterior position of the median pterygoid plate, and occlusion of the mandibular fenestra by a lateral plate of the dentary. Intriguingly, even in the other kingoriids which retain a pineal foramen (Dicynodontoides spp. and Kombuisia antarctica), this structure is reduced in size relative to other dicynodonts, suggesting that the pineal eye was less important for thermoregulatory activity in this clade than in other anomodonts. Although part of a local fauna including taxa that are otherwise widespread in the Karoo Basin (Daptocephalus, Lystrosaurus), the unique presence of Thliptosaurus in the relatively poorly-sampled Daptocephalus Assemblage Zone deposits of KwaZulu-Natal suggests that this region may preserve endemic taxa, and should be prioritized for future fieldwork.Item Rediscovery of the holotype of Clelandina major Broom, 1948 (Gorgonopsia: Rubidgeinae) with implications for the identity of this species(Evolutionary Studies Institute, 2017-12) Kammerer, Christian F.No specimen number was given for the holotype of the rubidgeine gorgonopsian species Clelandina major Broom, 1948 in its original description. Historically, a specimen in the Rubidge Collection (RC 94) was considered to represent Broom’s type specimen for C. major. However, recent study has revealed that the holotype of C. major is in fact a different specimen in the McGregor Museum in Kimberley (MMK 5031). The morphology of this specimen is consistent with the genus Clelandina, contra work based on RC 94 that considered C. major referable toAelurognathus. Clelandina major is here considered synonymous with the type species Clelandina rubidgei.MMK5031 represents only the fifth known specimen of this rare and unusual gorgonopsian.Item Two unrecognised burnetiamorph specimens from historic Karoo collections(2016-03) Kammerer, Christian F.Two historical specimens from Permian rocks of the Karoo Basin represent previously unrecognised members of the rare therapsid group Burnetiamorpha. These specimens cannot be referred to any existing burnetiamorph species, but are left in open nomenclature because of their incompleteness (both are isolated skull roofs). The first specimen is from the Tapinocephalus Assemblage Zone (AZ) and is characterized by heavily pachyostosed supraorbital bosses and a low nasal crest. The second specimen is from the Tropidostoma AZ and is generally similar to the Malawian taxon Lende, but is unique among described burnetiamorphs in having a frontoparietal ‘dome’ that surrounds the pineal foramen. Phylogenetic analysis of burnetiamorphs recovers support for a split between Proburnetia and Burnetia-like burnetiids, here named Proburnetiinae subfam. nov. and Burnetiinae Broom, 1923.