Palaeontologia africana

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ISSN (print): 0078-8554 ISSN (electronic): 2410-4418 For queries regarding content of Palaeontologia africana collections please contact Jonah Choiniere by email : jonah.choiniere@wits.ac.za or Tel : 011 717 6684

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    A new middle Permian burnetiamorph (Therapsida: Biarmosuchia) from the South African Karoo filling a gap in the biarmosuchian record
    (The Evolutionary Studies Institute, 2024-09) Matlhaga, Fonda; Benoit, Julien; Rubidge, Bruce S
    Biarmosuchia, the most basal therapsid clade, is represented by relatively few specimens known from Permian deposits in Russia and southern Africa. In both the Guadalupian (middle Permian) and Lopingian (late Permian), biarmosuchians represent less than 1% of the fossil record at the specimen level. Here, we describe a new burnetiamorph biarmosuchian, Impumlophantsi boonstrai, based on a partial skull and associated postcrania from the upper Tapinocephalus Assemblage Zone. It is characterized by the presence of a low nasal crest with a unique morphology among burnetiamorphs. Inclusion of this taxon in an updated phylogenetic analysis of biarmosuchians indicates that this specimen is one of the most basal burnetiamorphs, representing the only record of this grade from the middle Permian.
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    Observations on the structure of the Early Permian reptile Stereosternum tumidum Cope
    (Bernard Price Institute for Palaeontological Research, 1999) Modesto, Sean Patrick
    New information on the mesosaur Stereosternum tumidum is derived from a nearly complete skeleton and other material. Two autapomorphies are identified for Stereosternum: (1) presence of an ‘odontoid’ axial process, formed probably by co-ossification of the atlantal pleurocentrum to that of the axis, and (2) the presence of a posterior supraneural process on the neural arch of dorsal vertebrae. Temporal fenestration appears to be absent in Stereosternum, marginal teeth are determined to be subcircular rather than oval in cross section and in this respect resemble those of Mesosaurus, and there is no sign of fracture planes in the caudal vertebrae that could be indicative of caudal autotomy. A phylogenetic analysis, based on a slightly modified data matrix from the literature, identifies Mesosauridae as the sister group of Parareptilia within the reptilian clade Anapsida (sensu lato). As a consequence of this rearrangement of amniote tree topology, the stem-based taxon Sauropsida is regarded to be in abeyance, because it now includes exactly the same taxa as Reptilia. Mesosaurs, at more than 275 million years of age, can be recognised as the oldest known anapsid reptiles.
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    Preliminary phylogenetic analysis and stratigraphc congruence of the dicynodont anomodonts (Synapsida: Therapsida)
    (Bernard Price Institute for Palaeontological Research, 2001) Angielczyk, Kenneth D
    A preliminary phylogenetic analysis of 20 well-known dicynodont taxa was conducted using modern cladistic methods. Many past phylogenetic hypotheses were corroborated, but others conflict with the results of this analysis. Most notably, Diictodon, Robertia, and Pristerodon are reconstructed in more basal positions than previously suggested, whereas Endothiodon and Chelydontops occupy a more crown ward position. These findings are consistent with novel evolutionary scenarios for characters such as the presence of postcanine teeth and anterior palatal ridges. The Relative Completeness Index and Gap Excess Ratio were used to examine the degree of fit between the most parsimonious cladograms of this study and the stratigraphic record of the dicynodonts. Although the results of this analysis suggest that the preferred cladogram is relatively consistent with stratigraphy, the presence of some ghost ranges and ghost lineages imply that the fossil record of dicynodonts is not as complete as is sometimes stated. These findings are important because there is a long tradition of intensive collecting in regions where dicynodont fossils are common; sections of several dicynodont lineages may not be preserved in these areas.
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    Character state transformations and the fit of phylogenies to the fossil record
    (Bernard Price Institute for Palaeontological Research, 2002) Angielczyk, Kenneth D
    There is only one true history of life, and the biostratigraphic record and the phylogenetic relationships of organisms provide the most important information regarding this history. Ideally, the historical signal preserved in each of the data sets should be the same, and several methods have been proposed to compare the fit of phylogenies to the fossil record. All of these techniques use stratigraphic data associated with taxa, but our ability to recognize taxa and reconstruct their phylogenetic relationships ultimately is based on patterns of character state distributions that we observe. This raises the question of whether character states can be used to measure the fit of a phylogeny to the fossil record. Here I argue that we can, if the order of appearance of character states is considered. Optimization of character states on a phylogeny results in a predicted order of appearance of character states; derived states must arise after basal states. This order can be compared to that predicted by the fossil record. Although a number of factors can affect the frequency at which derived character states are sampled before basal states in the fossil record, conflicts between the two data sets should be relatively rare. Phylogenies that imply a large number of character state transformations that are inconsistent with the fossil record may need to be reconsidered before the fossil record is criticized.
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    The phylogeny of basal archosaurs
    (Bernard Price Institute for Palaeontological Research, 1994) Juul, Lars
    Archosaur systematics has received much attention from the mid 1970s and several influential works on this topic have emerged. As discrepancy exists among proposed phylogenies, some of the most important of the papers in question are assessed here. Characters used in cladistic analysis have often been selected too uncritically or phrased too vaguely to be of diagnostic value, and previously used, seemingly valid apomorphies have been disregarded by later workers, sometimes for no apparent reason. In the present paper a character matrix for the Archosauria was assembled by critically incorporating, and often modifying, characters used in earlier works on archosaur systematics. A phylogeny resulting from cladistic analysis of the matrix compiled here supports the monophyly of Archosauria, Crown-group Archosauria Crurotarsi, Ornithodira, Dinosauromorpha and Dinosauria, and disputes the existence of Suchia less Ornithosuchidae and of a monophyletic Crocodylotarsi. A new taxon, Dromaeosuchia, is erected for a clade consisting of Ornithosuchidae, Crocodylomorpha and Gracilisuchus plus Postosuchus.