Palaeontologia africana

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ISSN (print): 0078-8554 ISSN (electronic): 2410-4418 For queries regarding content of Palaeontologia africana collections please contact Jonah Choiniere by email : jonah.choiniere@wits.ac.za or Tel : 011 717 6684

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Start date: 2000End date: 2019

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    (DURASPACE, 2005-08-17) Diol, Joe
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    Pleistocene vertebrate trace fossils of Robberg Nature Reserve
    (Evolutionary Studies Institute, 2019-12) Helm, Charles W.; Cawthra, Hayley C.; Hattingh, Rudolph; Hattingh, Sinèad; McCrea, Richard T.; Thesen, Guy H. H.
    More than 140 Late Pleistocene trace fossil sites have been identified in aeolianites and lithified foreshore deposits along a 350 kilometre stretch of the Cape south coast in South Africa. Robberg Nature Reserve lies within this area and contains a zone of concentration of such tracksites, which complement the Pleistocene vertebrate body fossil record and assist in shedding light on the palaeoenvironment and palaeoecology of the Palaeo-Agulhas Plain. Ichnofossil sites unique to or of special significance within Robberg Nature Reserve include a substantial palaeosurface exposure that allows an estimate of track density, the best-preserved rhinoceros trackway identified to date, very well preserved artiodactyl tracks in the formof natural casts, small equid tracks, large elephant transmitted tracks, and well preserved sub-surface golden mole burrows with a burrow chamber. Aeolianite layers at Robberg have recently been dated to Marine Isotope Stage (MIS) 3. Samples that we have obtained for dating from the main palaeosurface underlie these dated layers and are anticipated to contribute to the understanding of the Pleistocene geology of the Robberg Peninsula. The protected status of the area lends itself to conservation, replication, interpretation and education initiatives.
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    The Younger Dryas interval at Wonderkrater (South Africa) in the context of a platinum anomaly
    (Evolutionary Studies Institute, 2019-10-02) Thackeray, J. Francis; Scott, Louis; Pieterse, P
    Wonderkrater in the Limpopo Province in South Africa is a late Quaternary archaeological site with peat deposits extending back more than 30 000 years before the present. Palaeoclimatic indices based on multivariate analysis of pollen spectra reflect a decline in temperature identifiable with the Younger Dryas (YD). A prominent spike in platinum is documented in aWonderkrater sample (5614) with a mean date of 12 744 cal yr BP using a Bayesian model, preceding the onset of the YD cooling event. The YD platinum spike at Wonderkrater is the first to be observed in Africa in the southern hemisphere, supplementing new discoveries from Patagonia in South America, in addition to more than 25 sites with such platinum anomalies in the northern hemisphere. The observations from South Africa serve to strengthen ongoing assessments of the controversial YD Impact Hypothesis, whereby it is proposed that a meteorite or cometary impact contributed to a decline in temperature, associated inter alia with dispersion of atmospheric dust, mammalian extinctions and cultural changes.
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    Cranial morphology and phylogenetic relationship of the enigmatic dinocephalian Styracocephalus platyrhynchus from the Karoo Supergroup, South Africa
    (Evolutionary Studies Institute, 2019-09) Fraser-King, Simon W.; Benoit, Julien; Day, Michael O.; Rubidge, Bruces S.
    Styracocephalus platyrhynchus is an unusual dinocephalian therapsid, known only from a handful of specimens from the Tapinocephalus Assemblage Zone of South Africa. It has had a chequered taxonomic history, largely because it is characterized by cranial pachyostosis and the presence of horn-like structures that project posteriorly from the temporal region; these features are found in the clades Burnetiamorpha and Dinocephalia. Its affinities have been further obfuscated by a lack of well-preserved material. This paper presents a description of a well-preserved skull referable to Styracocephalus from the western Karoo Basin and provides a revised generic diagnosis for the genus. This study – incorporating comparative anatomy,CT scanning, and cladistic analysis – reveals new character information that was not evident from pre-existing Styracocephalus material, and incorporates this into a new phylogenetic analysis. Our analysis recovers Styracocephalidae as a well-supported, monotypic family within Tapinocephalia, which is characterized by: prominent pachyostotic nasal and supraorbital bosses; two posteriorly projecting crest-like protuberances comprising contributions by the postorbital, squamosal and tabular bones; weak lingual heels on the incisor and postcanine dentition present with a moderate upper and lower canine. As Styracocephalus is restricted to the upper part of the Tapinocephalus Assemblage Zone, it may be a useful biostratigraphic index taxon in future.
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    Systematic experiments to quantitatively assess image quality for CT scans of a Karoo tetrapod fossil
    (Evolutionary Studies Institute, 2019-08) Tshibalanganda, Muofhe; du Plessis, Anton; Le Roux, Stephan G.; Taylor, Wendy L.; Smith, Roger M. H.; Browning, Claire
    Over the past decade non-destructive, three-dimensional visualization and analysis of fossils using X-ray tomography has greatly advanced palaeontological studies worldwide. Micro-computed tomography (microCT) is now accepted as best practice in palaeontological studies to augment the anatomical description of newly discovered fossils. Despite advances in laboratory microCT hardware, software and skills of users, there is a lack of clear methodologies for scanning and analysing fossils. Here we report on a systematic and detailed study of the quantitative effects of the variation of different microCT scanning parameters on the image quality of an unprepared fossilized Karoo tetrapod skull and parts of the postcrania. Results indicate that voltage variations do not increase the contrast for the bone as one would expect, and the best image quality solution is found using high frame averaging and high X-ray flux (current). Although this study was limited to one specimen, the results may find a practical use for future studies involving similar fossils.
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    First occurrence of the dicynodont Digalodon (Therapsida, Anomodontia) from the Lopingian upper Madumabisa Mudstone Formation, Luangwa Basin, Zambia
    (Evolutionary Studies Institute, 2019-04) Angielczyk, Kenneth D.
    Digalodon is a rare emydopoid dicynodont first described from upper Permian rocks in the Karoo Basin of South Africa. During fieldwork in the upper Madumabisa Mudstone Formation of the Luangwa Basin (Zambia) in 2014, a small dicynodont skull was discovered that conforms very well to the recently revised diagnosis of Digalodon rubidgei, although some minor differences between the Zambian and South African specimens are apparent. The Zambian occurrence of Digalodon expands the known geographic range of the genus, which was previously limited to a small set of localities in the vicinity of the town of Graaff-Reinet (Eastern Cape). Based on historical specimens, Digalodon is thought to have a comparatively short stratigraphic range in the Balfour Formation that spans the boundary between the Cistecephalus and Daptocephalus assemblage zones. This observation may allow refinement of biostratigraphic correlations between the Karoo and Luangwa Basins, but discovery of more precisely-provenanced specimens in the Karoo is needed to fully assess Digalodon’s biostratigraphic utility.
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    Postcranial osteology of the neotype specimen of Massospondylus carinatus Owen, 1854 (Dinosauria: Sauropodomorpha) from the upper Elliot formation of South Africa
    (Evolutionary Studies Institute, 2019-04) Barrett, Paul M.; Chapelle, Kimberley E.J.; Staunton, Casey K.; Botha, Jennifer; Choiniere, Jonah N.
    Massospondylus carinatus Owen, 1854, from the earliest Jurassic upper Elliot Formation of South Africa, was one of the first dinosaurs to be described from Gondwana. It has been incorporated into numerous phylogenetic, palaeobiological and biostratigraphic analyses, is often viewed as an exemplar for understanding sauropodomorph anatomy and is a key taxon in studies of early dinosaur evolution. Since its initial description, numerous specimens have been referred to this species, ranging from isolated postcranial elements to complete skeletons with three-dimensional skulls. In addition,M. carinatus has been identified in areas outside of the main Karoo Basin. Surprisingly, however, there have been few attempts to define the taxon rigorously, so that the basis for many of these referrals is weak, undermining the utility of this abundant material. Here, we provide the first detailed postcranial description of the neotype specimen ofM. carinatus, use it as a basis for diagnosing the species on the basis of cranial, axial and appendicular characters, demonstrate that it represents an adult individual on the basis of osteohistology, and discuss ways in which these data can assist in providing a better understanding of Karoo-aged African dinosaur faunas.
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    A fossil chrysochlorid skull in the Ditsong National Museum of Natural History: Robert Broom’s missing specimen unearthed?
    (Evolutionary Studies Institute, 2019-04) Mason, Matthew J.; Bennett, Nigel C.; Pickford, Martin
    An unlabelled, fossilized skull of a golden mole from the Ditsong National Museum of Natural History, Pretoria, was examined through micro-computed tomography. Reconstructions show that the species in question has alveoli for nine teeth in each upper jaw, although only six teeth in total remain. The skull resembles those of the extant chrysochlorids Amblysomus and Calcochloris, which also have nine teeth, but in most respects it is closer to Amblysomus. The ear region, examined in detail, also proved to be very similar to that of Amblysomus. Damage to the teeth and palate are consistent with the brief descriptions of a fossil golden mole skull first mentioned by Robert Broom in 1948. This specimen, dating from the Plio-Pleistocene and provisionally identified as Proamblysomus antiquus, subsequently went missing. We argue that the skull described here is Broom’s lost specimen, but whether this fossil species really deserves generic distinction from extant groups remains unclear.
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    The inner craniodental anatomy of the Papio specimen U.W. 88-886 from the Early Pleistocene site of Malapa, Gauteng, South Africa
    (Evolutionary Studies Institute, 2019-04) Bouchet, Florian; Ribéron, Alexandre; Heaton, Jason L.; Hoffman, Jakobus; Bam, Lunga; Jakata, Kudakwashe; Tawane, Mirriam; Tenailleau, Christophe; Zipfel, Bernhard; Beaudet, Amélie
    Cercopithecoids represent an essential component of the Plio-Pleistocene faunal assemblage. However, despite the abundance of the cercopithecoid fossil remains in African Plio-Pleistocene deposits, the chronological and geographic contexts from which the modern baboons (i.e. Papio hamadryas ssp.) emerged are still debated. The recently discovered Papio (hamadryas) angusticeps specimen (U.W. 88-886) from the Australopithecus sediba-bearing site of Malapa, Gauteng, South Africa, may represent the first modern baboon occurrence in the fossil record. Given the implication of U.W. 88-886 for the understanding of the papionin evolutionary history and the potential of internal craniodental structures for exploring evolutionary trends in fossil monkey taxa, we use X-ray microtomography to investigate the inner craniodental anatomy of this critical specimen. Our goal is to provide additional evidence to examine the origins of modern baboons. In particular,we explore (i) the tissue proportions and the dentine topographic distribution in dental roots and (ii) the endocranial organization. Consistent with the previous description and metrical analyses of its external cranial morphology, U.W. 88-886 shares internal craniodental anatomy similarities with Plio-Pleistocene and modern Papio, supporting its attribution to Papio (hamadryas) angusticeps. Interestingly, average dentine thickness and distribution in U.W. 88-886 fit more closely to the extinct Papio condition, while the sulcal pattern and relative dentine thickness are more like the extant Papio states. Besides providing additional evidence for characterizing South African fossil papionins, our study sheds new light on the polarity of inner craniodental features in the papionin lineage.
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    A new dicynodont (Anomodontia: Emydopoidea) from the terminal Permian of KwaZulu-Natal, South Africa
    (Evolutionary Studies Institute, 2019-04) Kammerer, Christian F.
    A new taxon of dicynodont (Thliptosaurus imperforatus gen. et sp. nov.) is described based on a dorsoventrally-crushed skull from latest Permian (upper Daptocephalus Assemblage Zone) strata in KwaZulu-Natal, South Africa. Thliptosaurus is distinguished from all other dicynodonts by an elongate intertemporal bar with broad dorsal exposure of the parietals but apparently no pineal foramen. Absence of the pineal foramen in dicynodonts is exceedingly rare; the only other taxa which exhibit this feature either have substantially broader (Kawingasaurus fossilis) or narrower (Kombuisia frerensis) intertemporal regions. Inclusion of Thliptosaurus in a phylogenetic analysis of dicynodonts recovers it as a kingoriid emydopoid, a position supported by its anteriorly-restricted pterygoid keel, elongate, curved anterior process of the lacrimal, relatively posterior position of the median pterygoid plate, and occlusion of the mandibular fenestra by a lateral plate of the dentary. Intriguingly, even in the other kingoriids which retain a pineal foramen (Dicynodontoides spp. and Kombuisia antarctica), this structure is reduced in size relative to other dicynodonts, suggesting that the pineal eye was less important for thermoregulatory activity in this clade than in other anomodonts. Although part of a local fauna including taxa that are otherwise widespread in the Karoo Basin (Daptocephalus, Lystrosaurus), the unique presence of Thliptosaurus in the relatively poorly-sampled Daptocephalus Assemblage Zone deposits of KwaZulu-Natal suggests that this region may preserve endemic taxa, and should be prioritized for future fieldwork.