Wits Evolutionary Studies Institute (ESI)
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Item Taxonomic and taphonomic interpretations of newly excavated in situ GD 2 faunal remains at Gondolin(The Evolutionary Studies Institute, 2024-01) Engelbrecht, Micke; Val, Aurore; Kibii, Job M.; Steininger, ChristineGondolin is one of the fossil-bearing karstic localities in the Cradle of Humankind, South Africa. Periodic excavations of calcified and decalcified sediments at several loci (GD 1,GD2 andGDA) over the last few decades have yielded a sizeable sample of Plio-Pleistocene fauna, including two hominin teeth from ex situ deposits. In 2015, renewed excavations were conducted at the GD 2 locality, which consists of decalcified in situ deposits, in order to shed more light on the site’s complex formation processes as well as to try finding new hominin material from a stratigraphically secured context. While these excavations did not yield any hominin material, abundant macrovertebrate remains were recovered. This paper presents the taxonomic composition and taphonomic characteristics of this new faunal assemblage. The occurrence of Equus sp. as well as the extinct species Metridiochoerus andrewsi and Hystrix makapanensis places the assemblage in a depositional age bracket of 2.33–1.78 Ma, which is consistent with ages already proposed forGD2. No primate material was recovered. The assemblage displays a taxonomic and bodyweight bias towards small-bodied (size classes I and II) bovids. The taphonomic characteristics of the bovid remains suggest the selective action of a leopard-like carnivore and while there is no direct evidence that the locality was used as a hyaena den, secondary scavenging by hyaenids cannot be excluded. Porcupines played an ancillary role in the bone accumulation. We performed intra- and inter-site taxonomic and taphonomic comparisons between this faunal sample and fossil assemblages from previously excavated localities at Gondolin (GD 1, GD 2 and GD A), as well as from other Paranthropus robustus-bearing sites in the Cradle of Humankind. These comparisons indicate that this new sample closely resembles the faunal assemblage previously collected from in situ calcified sediments at the GD 2 locality, in terms of species composition, bovid size class distribution, carnivore to ungulate ratio, and general taphonomic characterization.Item Renewed investigations at Taung; 90 years after the discovery of Australopithecus africanus(2016-10-18) Kuhn, Brian F; Herries, Andy I R; Price, Gilbert J; Baker, Stephanie E; Hopley, Philip; Menter, Colin; Caruana, Matthew V2015 marked the 90th anniversary of the description of the first fossil ofAustralopithecus africanus, commonly known as the Taung Child, which was unearthed during blasting at the Buxton-Norlim Limeworks (referred to as the BNL) 15 km SE of the town of Taung, South Africa. Subsequently, this site has been recognized as a UNESCOWorld Heritage site on the basis of its importance to southern African palaeoanthropology. Some other sites such as Equus Cave and Black Earth Cave have also been investigated; but the latter not since the 1940s. These sites indicate that the complex of palaeontological and archaeological localities at the BNL preserve a time sequence spanning the Pliocene to the Holocene. The relationship of these various sites and how they fit into the sequence of formation of tufa, landscapes and caves at the limeworks have also not been investigated or discussed in detail since Peabody’s efforts in the 1940s. In this contribution we mark the 90th anniversary of the discovery and description of the Taung Child by providing a critical review of previous work at Taung based on our recent preliminary work at the site. This includes a reassessment of the Taung Child Type Site, as well as renewed excavations at Equus Cave and the lesser-known locality and little-investigated Black Earth Cave. Preliminary results suggest that much of our previous understandings of the BNL’s formational history and site formation processes need to be reassessed. Only through detailed analysis on the BNLas a whole can we understand this complex depositional environment.Item The macrovertebrate fossil assemblage from the Name Chamber, Sterkfontein: taxonomy, taphonomy and implications for site formation processes(2015-11-27) Val, Aurore; Stratford, Dominic JThe Name Chamber contains some of the deepest fossiliferous deposits in the Sterkfontein Caves, at ~20 metres below the surface deposits of Members 4 and 5. Recent excavations complemented by detailed studies of the geological context, as well as of the microfaunal and lithic (i.e. Oldowan artefacts) assemblages, have shed some light on the complex history of sediment accumulation in that part of the Sterkfontein karstic system. The Name Chamber has a long and intricate history of deposition that is of particular value for understanding the redistribution of Oldowan-bearing sediments from Member 5 into the deep chamber below. Recognizing sediment movement and sources through multidisciplinary investigations is of key importance to reconstructing primary lithic assemblages and ultimately the behavioural proxies associated with them. Here, we present the results of a taxonomic and taphonomic analysis of the macrofaunal assemblage recovered during excavations of the decalcified sediments of the Western Talus in the Name Chamber. The taxonomic composition of the faunal spectrum is similar to that of Member 5 East Oldowan, with an overrepresentation of medium-sized bovids and the occurrence of taxa associated with grasslands. The taphonomic features of the fossil remains are characteristic of a mixed assemblage with indications of contributions by carnivores, slope wash and gravity collecting bones from the catchment surface (including carnivore-gnawed and butchery-marked specimens). The results independently corroborate lithic and microfaunal analyses and support the hypothesis of multiple origins for the sediments of the Name Chamber, with a main contribution from Member 5 East Oldowan (notably illustrated in the Name Chamber assemblage by the identification of several cut-marked remains and a bone tool), shortly after it accumulated at about 2.18 Ma. There is also indication of a minor contribution from Member 4 but no evidence for a noticeable contribution from Post-Member 6 (L/63 Infill).Item Superior cervical vertebrae of a Miocene hominoid and a Plio-Pleistocene hominid from southern Africa(Bernard Price Institute for Palaeontological Research, 2000) Gommery, DominiqueThe Miocene hominoid and Plio-Pleistocene hominid vertebral record is poor. In 1994, a complete atlas of a hominoid was found in breccia at Berg Aukas in Namibia. Its age was estimated to be middle Miocene (13 myr) on the basis of microfauna. This locality yielded the holotype of Otavipithecus namibiensis and the atlas could belong to the same genus. The specimen exhibits clear hominoid traits such as a weakly salient retroglenoid tubercle at the superior articular facet of the lateral mass, and a horizontal transverse process. This morphology of the transverse process is close to that of pygmy chimpanzees, gibbons and African colobines, suggesting that Otavipithecus was arboreal. This confirms the conclusions drawn from other parts of the skeleton. From the size of the atlas, a body weight of 15-20 kg is estimated for the Berg Aukas hominoid, which accords with previous estimates based on its teeth. The second fossil considered in this paper is an axis from Swartkrans, SK 854, dated to nearly 1,8 myr. This axis is compared with another Plio-Pleistocene axis from Ethiopia, AL 333.101. SK 854 shows a morphology different from that of humans and AL 333.101, and also of apes. The South African axis was attributed to Paranthropus by Robinson (1972), and its morphology is probably typical of bipedalism associated with climbing.Item New evidence of the Giant Hyaena, Pachycrocuta brevirostris (Carnivora, Hyaenidae), from the Gladysvale Cave deposit (Plio-pleistocene, John Nash Nature Reserve, Gauteng, South Africa)(Bernard Price Institute for Palaeontological Research, 2001) Mutter, Raoul J; Berger, Lee R; Schmid, PeterA well preserved cranium which represents the most complete skull of Pachycrocuta brevirostris (Carnivora, Hyaenidae) discovered in Africa, and a maxillary fragment from the Gladysvale Cave Deposit (John Nash Nature Reserve, Gauteng, South Africa) are described and compared to other fossil and extant hyaenid specimens from South Africa and Europe. In addition, some aspects of functional morphology in the hyaenid dentition are reconsidered and suggested to be directly related to the palaeoecological role of P. brevirostris.Item Locomotor and habitat classifications of cercopithecoid postcranial material from Sterkfontein Member 4, Bolt's Farm and Swartkrans Members 1 and 2, South Africa(Bernard Price Institute for Palaeontological Research, 2001) Elton, SarahMuch southern African cercopithecoid postcranial material is not associated with craniodental remains. Consequently, this postcranial material cannot be confidently assigned to a particular taxon, with the result that little is known about the locomotor strategies and habitat preferences of specific Plio-Pleistocene cercopithecoids from southern Africa. However, cercopithecoid postcrania can provide important information about habitats that were present at fossil sites, even when the material is not attributed to taxa. In this paper, ecomorphic analysis is used to assign cercopithecoid postcranial material from Sterkfontein Member 4, Bolt's Farm and Swartkrans Members 1 and 2 to one of three habitat/locomotor categories: forest-living arboreal quadrupeds (‘forest arboreal'), open habitat terrestrial quadrupeds (‘open terrestrial') and open habitat cercopithecoids using a mix of arboreal and terrestrial quadrupedalism (‘open mixed'). Cercopithecoids representing all three habitat categories were found in the samples from Sterkfontein Member 4 and Bolt' s Farm, suggesting that monkeys using a range of habitats and locomotor strategies were present at these sites. However, no 'forest arboreal' cercopithecoids were found in the samples from Swartkrans Members 1 and 2, indicating that cercopithecoids at these localities probably depended largely on open habitats. The habitat and locomotor strategy data were also used in combination with locality-based listings of fossil cercopithecoid craniodental remains to suggest possible locomotor strategies for several southern African Plio-Pleistocene cercopithecoid taxa, including Parapapio broomi (possibly 'forest arboreal'), Parapapio jonesi (‘open terrestrial'), Papio robinsoni (‘open terrestrial') and Cercopithecoides williamsi (‘open mixed').Item Undescribed suid remains from Bolt's Farm and other Transvaal cave deposits(Bernard Price Institute for Palaeontological Research, 1993) Cooke, H B SAlthough individual suid specimens from other sites have been described, only Makapansgat has been treated comprehensively, but much new material has come to light in the past three decades. The University of California African Expedition worked several sites of different ages at Bolt's Farm and the material recovered includes a virtually complete, slightly compressed, skull of Phacochoerus modestus (= P. antiquus Broom 1948) from Pit 3, associated with Antidorcas recki, and also several cranial and dental fragments from Pit 14 that belong to the typical Makapansgat Potamochoeroides shawi. Bolt's Farm was the source of a cranial specimen described by Broom as "Notochoerus meadowsi" (= Metridiochoerus andrewsi); other specimens referred to this taxon and described by Shaw as from "Sterkfontein Lime Works" more probably came from Bolt's Farm as well. Broom's cranium and a pair of mandibles have closer resemblances to Metridiochoerus jacksoni of East Africa. Swartkrans has yielded both described and undescribed material referred to Phacochoerus modestus and to Metridiochoerus. Discounting Shaw's material, only one small specimen has come from the Sterkfontein Type Site, a mandible fragment of a juvenile with an incompletely formed third molar in alveolus but it can be matched remarkably closely with a specimen from Makapansgat and there is very little doubt that it belongs to Potamochoeroides shawi. An undescribed third molar from the pink breccia at Makapansgat is comparable with early Notochoerus scotti from East Africa. The status of Notochoerus capensis is reconsidered.Item Fossil proboscidean remains from Bolt's Farm and other Transvaal cave breccias(Bernard Price Institute for Palaeontological Research, 1993) Cooke, H B SProboscidean remains are very rare in the Transvaal cave breccias and the few specimens recovered are generally fragmentary but deserve description because of their potential value in correlation and dating. The best specimen is the back half of a left lower molar from Pit 7 of the University of California African Expedition's work at Bolt's Farm. It was regarded by Maglio as representing Elephas ekorensis but closer examination suggests that there are some more progressive characters and it most likely represents Elephas recki brumpti. The stump of a second molar from Bolt's farm also accords with this taxon. Makapansgat has furnished 14 fragmentary fossils, 6 of them tusk or root remains. Particularly interesting is the occurrence of a pair of cones from a molar of Anancus. The scrappy elephantid material from Makapansgat may be referred tentatively to an early stage of the Elephas recki lineage, as also a mandible fragment from Sterkfontein with the two anterior milk teeth. An anterior milk tooth from Swartkrans Member 3 exhibits broader and higher lamellae than in E. recki and most probably belongs to the more advanced E. iolensis to which most of the Vaal River elephants have been referred.Item Taxonomic description of fossil wood from Cainozoic Sak River terraces, near Brandvlei, Bushmanland, South Africa(Bernard Price Institute for Palaeontological Research, 1993) Bamford, Marion K; de Wit, Michiel C JSeven pieces of silicified wood are described from two sites near the Sak River, Bushmanland. The Miocene deposit yielded five specimens which can be assigned to the Dipterocarpaceae, Fagaceae, Myrtaceae, Oleaceae and Rutaceae. Of the two logs recovered from the Plio-Pleistocene deposit, only one was well enough preserved to be assigned to the Polygalaceae. All the woods indicate that the palaeoenvironment in that region was tropical to subtropical based on the wood structure, growth rings and from their modem counterparts.