Wits Evolutionary Studies Institute (ESI)

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    A multi-disciplinary review of late Quaternary palaeoclimates and environments for Lesotho
    (2016-07) Fitchett, J.M.; Grab, S.W; Bamford, M.K.; Mackay, A.W.
    Lesotho provides a unique context for palaeoclimatic research. The small country is entirely landlocked by South Africa, yet has considerable variation in topography, climate, and associated vegetation over an approximate east–west transect. The region has been of archaeological interest for over a century, and hosts many Early to Late Stone Age sites with occupation preceding 80 000 years before present. The eastern Lesotho highlands are of interest to periglacial and glacial geomorphologists because of their well-preserved relict landforms and contentious evidence for permafrost and niche glaciation during the late Quaternary. However, continuous proxy records for palaeoenvironmental reconstructions for Lesotho are scarce and hampered by a range of methodological shortfalls. These challenges include uncertain ages, poor sampling resolution, and proxies extracted from archaeological excavations for which there may be bias in selection. Inferences on palaeoclimates are thus based predominantly on archaeological and palaeogeomorphological evidence for discrete periods during the late Quaternary. This review paper presents a more detailed multidisciplinary synthesis of late Quaternary conditions in Lesotho. We simultaneously considered the varying data that contribute to the under-studied palaeoenvironmental record for southern Africa. The collective palaeoenvironmental data for eastern Lesotho were shown to be relatively contradictory, with considerable variations in contemporaneous palaeoclimatic conditions within the study area. We argue that although methodological challenges may contribute to this variation, the marked changes in topography result in contrasting late Quaternary palaeoenvironments. Such environments are characterised by similar contrasting microclimates and niche ecologies as are witnessed in the contemporary landscape. These spatial variations within a relatively small landlocked country are of importance in understanding broader southern African palaeoenvironmental change.
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    A review of the reptile and amphibian assemblages from the Stormberg of southern Africa, with special emphasis on the footprints and the age of the Stormberg
    (Bernard Price Institute for Palaeontological Research, 1984) Olsen, Paul E.; Galton, Peter M.
    The Molteno, Elliot, and Clarens formations comprise the continental Stormberg Group of the Karoo Basin of South Africa and Lesotho. The Molteno Formation contains a well preserved macro- and microfloral assemblage but apparently no vertebrates; the Elliot and Clarens formations contain abundant vertebrates but virtually no floral remains. The vertebrate taxa represented by skeletal remains are listed and divided into two assemblages - the lower Stormberg (lower Elliot) and upper Stormberg (upper Elliot and Clarens) assemblages. The abundant, diagnosable footprint taxa are revised and their names reduced to eight genera. These ichnotaxa also fall into two biostratigraphic zones that parallel the skeletal assemblages. Comparison of the faunal assemblages with those of the European type section strongly suggests that the lower Stormberg assemblage is Late Triassic (Carnian- Norian) in age while the upper Stormberg assemblage is Early Jurassic (Hettangian-Pliens- bachian) in age. Comparisons with other continental assemblages from other areas suggest that the upper Stormberg (upper Elliot and Clarens formations) assemblage broadly correlates with the upper Newark Supergroup of eastern North America, the Glen Canyon of the southwestern United States, and the lower Lufeng Series of China- all thought to be of Early Jurassic age on the basis of floral and/or radiometric evidence. Based on these correlations, previously published paleobiogeographic maps are revised; these show a shift from Late Triassic floral and faunal provinciality to Early Jurassic homogeneity. This shift was synchronous with a widening of the equatorial arid zone.
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    Late Triassic traversodont cynodonts from Nova Scotia and southern Africa
    (Bernard Price Institute for Palaeontological Research, 1984) Hopson, James A.
    The first gomphodont cynodont from North America is described from the Upper Triassic Wolfville Formation, Fundy Group, Newark Supergroup, of Burntcoat, Minas Basin, Nova Scotia, Canada. Known material consists of a large mandible, edentulous but for two incisors, a probably associated canine, and two small dentaries; an isolated multicusped tooth may belong to this species. This gomphodont closely resembles the large traversodont Scalenodontoides macrodontes from the lower Elliot Formation (= Red Beds) of Lesotho; it is provisionally placed in this genus but is a distinct species, ?Scaleno- dontoides plemmyridon sp. nov. It differs from S. macrodontes primarily in its more massive symphyseal region and much larger mental foramen from which a prominent groove extends posterodorsally. The large, posteriorly-located mental foramen is believed to be a well-developed oral vestibule and cheek. The isolated tooth, provisionally interpreted as a traversodont lower postcanine, is anteroposteriorly compressed, with a high anterior blade formed by three transversely-aligned cusps and a short heel; it does not resemble postcanines of S. macrodontes and so reference to ?S. plemmyridon is questionable. Scalenodontoides is the sister genus of Exaeretodon from the Late Triassic of Argentina, Brazil, and India; they are allied on the basis of: upper incisors reduced from 4 to 3, all incisors greatly enlarged, and internarial bar incomplete. These resemblances to tritylodontids are convergent. Scalenodontoides and Exaeretodon share with Gomphodontosuchus the enlargement of the anterolabial cusp of the lower postcanines and posterior inclination of the anterolingual cusp. The Wolfville and basal Elliot faunas are considered to be Late Carnian or Carno-Norian in age. Faunas containing Exaeretodon are older Carnian, though the Santa Maria Formation of Brazil may be Late Ladinian.