Wits Evolutionary Studies Institute (ESI)

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Start date: 2000Subject: search.filters.subject.biostratigraphy

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    A new middle Permian burnetiamorph (Therapsida: Biarmosuchia) from the South African Karoo filling a gap in the biarmosuchian record
    (The Evolutionary Studies Institute, 2024-09) Matlhaga, Fonda; Benoit, Julien; Rubidge, Bruce S
    Biarmosuchia, the most basal therapsid clade, is represented by relatively few specimens known from Permian deposits in Russia and southern Africa. In both the Guadalupian (middle Permian) and Lopingian (late Permian), biarmosuchians represent less than 1% of the fossil record at the specimen level. Here, we describe a new burnetiamorph biarmosuchian, Impumlophantsi boonstrai, based on a partial skull and associated postcrania from the upper Tapinocephalus Assemblage Zone. It is characterized by the presence of a low nasal crest with a unique morphology among burnetiamorphs. Inclusion of this taxon in an updated phylogenetic analysis of biarmosuchians indicates that this specimen is one of the most basal burnetiamorphs, representing the only record of this grade from the middle Permian.
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    Re-identification and updated stratigraphic context of the holotypes of the late Permian tetrapods Dicynodon ingens and Scymnosaurus warreni from KwaZulu-Natal
    (Evolutionary Studies Institute, 2023) Groenewald, David P; Kammerer, Christian F
    Two historical therapsid holotypes held in the collections of the KwaZulu-Natal Museum (Dicynodon ingens Broom, 1907 and Scymnosaurus warreni Broom, 1907) are redescribed. The holotype of D. ingens is a snout tip of a large dicynodontoid that can be re-identified as a specimen of Daptocephalus leoniceps based on premaxillary proportions and palatal morphology. The holotype of S. warreni consists of a dorsoventrally crushed therocephalian snout, which is here re-identified as that of Moschorhinus kitchingi based on size, general proportions, tooth count, and septomaxillary morphology. The localities of both specimens are biostratigraphically assigned to the Lystrosaurus maccaigi-Moschorhinus Subzone of the Daptocephalus Assemblage Zone (latest Permian).
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    The first record of late Permian tetrapods from Namibia
    (Evolutionary Studies Institute, 2023) Mocke, Helke B; Kammerer, Christian F; Smith, Roger M H; Marsicano, Claudia A
    The entire Omingonde Formation in Namibia was previously assigned a Triassic age based on the fossil tetrapod fauna discovered southwest of theWaterberg Plateau. However, recently repatriated and newly collected material suggests that the lowermost part of the Omingonde Formation includes late Permian strata, equivalent in age to the upper Endothiodon Assemblage Zone of the South African Karoo Basin. In this study,we document the first record of late Permian tetrapod fossils from Namibia, and provide brief descriptions of a gorgonopsian skull collected at Mount Etjo in 1996 and the skull of a small dicynodont collected at the same site in 2019, referable to cf. Tropidostoma. This material confirms that there is need for better understanding of the stratigraphy of the Etjo Mountain area, in particular the position and nature of the unconformity between strata of the Permian Endothiodon and the Triassic Cynognathus assemblage zones.
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    New ‘rauisuchian’ fossil material from the lower Elliot Formation of South Africa
    (Evolutionary Studies Institute, 2023) Tolchard, Frederick B; Bordy, Emese M; Choiniere, Jonah N
    ‘Rauisuchians’ are a grade of paracrocodylomorph archosaurs that evolved a diversity of body plans and played a key role in ecosystems worldwide throughout the Triassic. They are widely believed to have gone extinct during the end-Triassic mass extinction event though the fossil record of rauisuchians in the latest Triassic is still poorly known. In this study, we describe new rauisuchian fossil remains from the lower Elliot Formation of South Africa. Based on comparative anatomical evidence, we assign these specimens to the pseudosuchian clade Rauisuchidae. The addition of this material to the existing southern African fossil record allows us, along with some new insights into the taxonomic affinities of previously published material, to identify the presence of at least three distinct taxa of rauisuchian in the Norian of southern Africa: two of the clade Rauisuchidae; and one of the clade Poposauroidea. These likely filled the ecological role of apex predators in the Late Triassic of southern Africa. We also provide a revision of the stratigraphic record of ‘rauisuchians’ from the Elliot Formation and show that the existing record of rauisuchians in southern Africa, though among the youngest in the world, does not extend beyond the late Norian. Further exploration of the Elliot Formation and other terrestrial deposits at the Triassic–Jurassic boundary will be needed to assess whether rauisuchians survived until, or even beyond, the end-Triassic mass extinction event.
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    First occurrence of the dicynodont Digalodon (Therapsida, Anomodontia) from the Lopingian upper Madumabisa Mudstone Formation, Luangwa Basin, Zambia
    (Evolutionary Studies Institute, 2019-04) Angielczyk, Kenneth D.
    Digalodon is a rare emydopoid dicynodont first described from upper Permian rocks in the Karoo Basin of South Africa. During fieldwork in the upper Madumabisa Mudstone Formation of the Luangwa Basin (Zambia) in 2014, a small dicynodont skull was discovered that conforms very well to the recently revised diagnosis of Digalodon rubidgei, although some minor differences between the Zambian and South African specimens are apparent. The Zambian occurrence of Digalodon expands the known geographic range of the genus, which was previously limited to a small set of localities in the vicinity of the town of Graaff-Reinet (Eastern Cape). Based on historical specimens, Digalodon is thought to have a comparatively short stratigraphic range in the Balfour Formation that spans the boundary between the Cistecephalus and Daptocephalus assemblage zones. This observation may allow refinement of biostratigraphic correlations between the Karoo and Luangwa Basins, but discovery of more precisely-provenanced specimens in the Karoo is needed to fully assess Digalodon’s biostratigraphic utility.
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    Biesiespoort revisited: a case study on the relationship between tetrapod assemblage zones and Beaufort lithostratigraphy south of Victoria West
    (Evolutionary Studies Institute, 2018-12) Day, Michael O.; Rubidge, Bruce S.
    The relationship between the tetrapod assemblage zones of the South African Karoo Basin and the lithostratigraphic divisions of the Beaufort Group is well-established, and provides an independent means of dating fossil occurrences. However, this relationship may not be consistent across the basin; a discrepancy exists between the historical tetrapod assemblages in the vicinity of Victoria West, Northern Cape Province, and the expected tetrapod assemblage zones based on mapped geology. In order to examine this disconnect, we collected fossils at two localities close to Biesiespoort railway station, a locality that was visited on a number of occasions by Robert Broom. Our fossil samples support the biostratigraphic determinations of Broom and thus confirmthat the stratigraphic extent of the biozones at these localities differs from their type areas further south. The reasons for this are unclear but could be related to the northward younging of the lithological units, implying complex depositional processes, or result from difficulties in mapping. Nevertheless, caution should be exercised when using mapped geology near Victoria West as a guide to the age of fossils found there.
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    A new GIS-based biozone map of the Beaufort Group (Karoo Supergroup), South Africa
    (Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, 2010-12) van der Walt, Merrill; Day, Michael; Rubidge, Bruce; Cooper, Antony K.; Netterberg, Inge
    The stratigraphic subdivision of the Karoo Supergroup of South Africa has been addressed for more than a century by various geoscience studies. A lack of good lithostratigraphic markers in the Beaufort Group and the fact that the succession preserves a rich diversity of therapsid fossils for which there is a robust taxonomic scheme, make biostratigraphy the best option for correlation. This is applicable both within the basin and when comparing the Karoo succession to other coeval depositional basins. Published biostratigraphic maps of the Beaufort Group were compiled based on rough estimates of the distribution of zone-defining tetrapod fossil genera throughout the basin. None of these simultaneously utilized all the databases of South African museum collections. The recent application of GIS technology to integrate the vertebrate fossil databases of all South African museums has resulted in a far more precise biozone map which can be continuously updated as new information is received. This digital map introduces an entirely new way of representing the geographical distribution of fossil data, and thus can potentially enhance basin development and tetrapod biogeographic studies.
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    A second specimen of Blikanasaurus (Dinosauria: Sauropoda) and the biostratigraphy of the lower Elliot Formation
    (Bernard Price Institute for Palaeontological Research, university of the Witwatersrand, 2008-04) Yates, Adam M.
    A second specimen of the rare basal sauropod Blikanasaurus cromptoni, is recorded from a site in the Ladybrand district of the Eastern Free State, South Africa. The specimen consists of a right metatarsal 1 that originated from the upper 20mof the lower Elliot Formation. It can be referred to B. cromptoni on the basis of its small size and highly robust proportions, which distinguish this taxon from all other sauropodomorphs. This record extends the geographic distribution of B. cromptoni north into the region of the main Karoo Basin where the Elliot Formation is dramatically thinner. It also extends the known stratigraphic range of B. cromptoni up from the base of the Elliot Formation into a position near the top of the lower member. This new record, combined with other new discoveries, supports the hypothesis that the thin northern part of the lower Elliot Formation is a condensed section that is largely, if not entirely, coeval with the thicker southern sections.
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    A new cynodont record from the Tropidostoma Assemblage Zone of the Beaufort Group: implications for the early evolution of cynodonts in South Africa
    (Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, 2008-04) Botha-Brink, J.; Abdala, F.
    A new specimen of cynodont has been recovered from the lower Upper Permian levels of the Tropidostoma Assemblage Zone of the South African Karoo Basin. A series of characters, including the presence of an interpterygoid opening, triconodont postcanine teeth, which are circular in crown view, a probable lingual cingulum, and most significantly, the presence of a masseteric fossa high on the coronoid process, allows this specimen to be assigned to the Procynosuchidae. However, unlike most procynosuchids, precanine teeth are absent and the incisors are represented by four left and five right upper and probably, three lower teeth. Considering the small size of the new specimen from the Tropidostoma Assemblage Zone we identify it as a juvenile Procynosuchus delaharpeae, the only species currently recognized in the genus. The low number of incisors and absence of precanines in this specimen are interpreted as ontogenetic variation (i.e. juvenile characters). This discovery extends the biostratigraphic range of Procynosuchus, as it is now recorded in three assemblage zones, indicating that this taxon is the longest-lived cynodont in the Karoo Basin. The new specimen of Procynosuchus and the recently described Charassognathus gracilis, are the oldest global records of cynodonts, and indicate an earlier initial radiation of this lineage than was previously thought.
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    Charting the fossils of the Great Karoo: a history of tetrapod biostratigraphy in the Lower Beaufort Group, South Africa
    (Evolutionary Studies Institute, University of the Witwatersrand, 2013-12-18) Day, Mike
    The interest in the fossil remains of the Beaufort Group and their stratigraphic significance goes back as far as the earliest geological studies in South Africa in the early 19th century. By the 1890s, the understanding of fossil distributions in the sedimentary rocks of the Karoo allowed the formulation of the first tetrapod biostratigraphic subdivisions. Since the beginning of the 20th century, the highest resolution subdivisions of the mostly undifferentiated fluvial sediments of the Beaufort Group have been biostratigraphic. More recent biostratigraphic studies in the Lower Beaufort Group have been crucial in understanding terrestrial ecological change in the Middle and Late Permian, and continue to be a leading area of research in South Africa palaeontology.