Wits Evolutionary Studies Institute (ESI)

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    A problematical element in the Glossopteris flora of Vereeniging
    (Bernard Price Institute for Palaeontological Research, 1975) LeRoux, S F
    An unusual element, not previously recorded from the mixed Glossopteris Flora of Vereeniging, is described here. The type material was obtained from a rich fossiliferous zone, previously described by the author (Le Roux, 1963, pp. 1-2). The material available for study consists of a single specimen of an incomplete frond preserved in the form of an impression in fine-grained laminated clay.
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    The morphology and relationships of Youngina capensis Broom and Prolacerta broomi Parrington
    (Bernard Price Institute for Palaeontological Research, 1975) Gow, Chris E
    Comprehensive descriptions of the osteology of Youngina capensis Broom and Prolacerta broomi Parrington are presented. New details of the braincase of Proterosuchus fergusi Broom are given as these became necessary for comparative purposes. It is suggested that the initial radiation of sauropsid reptiles was a Permian event as yet poorly documented. The phylogenetic position of Youngina forward and backward in time cannot be narrowly defined, though certain characters seem specifically to preclude it from lizard ancestry. Prolacerta, on the basis of tooth implantation, braincase morphology and postcranial anatomy is shown to be closest to the proterosuchian thecodonts. It is very definitely not concerned with lizard origins, but would on available evidence seem to be a perfectly good ancestor for the middle Triassic forms Macrocnemus and Tanystropheus which latter must cease to be regarded as lizard ancestors. We have here a rather distinct reptilian lineage which branched off from common ancestral stock just prior to the advent of archosaurs.
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    The affinities of Proterochampsa barrioneuvoi Reig
    (Bernard Price Institute for Palaeontological Research, 1975) Cruickshank, Arthur R I
    Proterochampsa barrioneuvoi Reig is re-examined and is confirmed as a proterosuchian thecodont. None of the features previously thought to ally it to the Crocodilia are solely characteristic of that group. On the other hand it is not a phytosaur nor phytosaur ancestor, only showing one real trend towards these animals in the rearward migration of the internal and external nares. Proterochampsa and its relatives Chanaresuchus, Gualosuchus and Cerritosaurus are too late in time to be phytosaur ancestors. They are grouped together in the Proterochampsidae, a family within the Proterosuchia.
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    Fossil wood of Cretaceous age from the Namaqualand continental shelf, South Africa
    (Bernard Price Institute for Palaeontological Research, 1994) Bamford, Marion K; Corbett, Ian B
    Fossil wood was collected from sediments on the Namaqualand shelf, West Coast, South Africa, between the Orange River mouth and just to the south of Kleinzee. Forty three of these samples are here described. All the woods are gymnospermous and have abietinian tracheid pitting. Nineteen of them are well enough preserved to be identified to species level: Podocarpoxylon cf. umzambense, Mesembrioxylon cf. stokesii, M. cf. sahnii, M. cf. woburnense and Protocupressinoxylon cf. purbeckensis. The remainder of the woods have been placed in the artificial genus Mesembrioxylon without species names. The woods are probably primitive members of the Podocarpaceae growing during the Lower Cretaceous. They indicate a seasonal climate and inhabited the extensive low-lying coastal regions.
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    The stratigraphy of the Sterkfontein hominid deposit and its relationship to the underground cave system
    (Bernard Price Institute for Palaeontological Research, 1991) Partridge, T C; Watt, I B
    A programme of orientated core drilling was carried out during 1989 to elucidate stratigraphic relationships within the Sterkfontein Formation and to obtain a representative suite of samples for palaeomagnetic analysis. The cores have revealed that the hominid-bearing cave deposits occur as a continuous succession comprising 6 Members and extending to a maximum depth of about 30 m below present surface. Of these Member I (comprising a sterile, residual fill) and Member 3 are the most extensive. This sequence has been displaced vertically downwards within a zone of decalcification coinciding with the central part of the deposit. This zone has been the focus of recent deep excavations at the site. The results of the drilling, in conjunction with recent surveys of the underground cave system, confirm that a dolomite floor existed at an average depth of about 20 m at the time offirst cave filling. Subsequent cavern development down to depths in excess of 50 m caused the local collapse of some lower units of the Sterkfontein Formation and, as new openings developed to the surface, permitted the ingress of younger fills below the base of the hominid-bearing succession.
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    Palaeo-ecology of the Sterkfontein hominids: a review and synthesis
    (Bernard Price Institute for Palaeontological Research, 1991) McKee, Jeffrey K
    Excavations at the Sterkfontein hominid fossil site have yielded a rich and revealing faunal assemblage. Evolutionary transitions are evident in early hominids and associated fauna between the times represented by Members 4 and 5. Member 4 has yielded a large and variable sample of Australopithecus africanus as well as evidence of considerable species diversity among the artiodactyls, carnivores and primates. The appearance of early Homo along with stone and bone tools in Member 5 coincides with a reduction of species representation in the orders of larger mammals as well as with the occurrence of new derived species and apparent extinctions. Three hypotheses have been suggested to account for the trends seen in the hominid-bearing members of the Sterkfontein Formation. The 'Climatic Change Hypothesis' accounts for the evolutionary trends by the causal factors of global and local cooling and aridification with evidence of savanna-grasslands supplanting an earlier environment with a denser cover of vegetation. The 'Taphonomic Hypothesis' explains changes in relative species representation at Sterkfontein in terms of the bone-accumulating agents; in Member 4 primary carnivores were largely responsible for the deposition of large mammalian fauna, whereas the scavenging activities of early Homo would have accounted for much of the bone and all of the artefacts found in Member 5. A third proposition is the 'Species Interaction Hypothesis', a derivative of the ' Red-Queen Hypothesis'; here the dynamics of species interaction, including competition and commensalism among hominids, carnivores and cercopithecids, propel the evolutionary changes and cause the extinctions. These hypotheses are not mutually exclusive, but the relative effects of the factors involved must be verified or refuted with better chronological controls and further analyses of the African fossil sites. The Sterkfontein Formation represents a microcosm in which various scenarios of African mammalian evolution can be tested.
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    The palaeontology of Haasgat a preliminary account
    (Bernard Price Institute for Palaeontological Research, 1991) Keyser, Andre W
    Haasgat is a cave on the steep western slope of the upper reach of the Witwatersrand Spruit, on the farm Leeuwenkloof 480 lQ, in the Brits District. It was heavily mined for flowstone (calcite). The cave contains a deposit offossiliferous cave silt and breccia that was partially removed by the miners and dumped on the steep slopes of the valley. The original entrance was probably a shallow inclined pit, leading into an upper chamber and then into the preserved depository. Both porcupines and carnivores served as accumulating agents for the bones. Fossils of the primates Parapapio and Cercopifhecoides, hyaena (Chasmaporthetes), fox, porcupines, several species of bovids and two species of Hyrax have been recovered. An insufficient number of fossils have been prepared to determine the age of the deposit with certainty. The deposit was provisionally thought to be of Pliocene age because of the occurrence of Parapapio. At this stage it would be unwise to correlate this occurrence with any other caves in this age range. It is concluded that the cave silts were deposited by flash floods, under a wetter climatic regime than that of the present.
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    A review of the reptile and amphibian assemblages from the Stormberg of southern Africa, with special emphasis on the footprints and the age of the Stormberg
    (Bernard Price Institute for Palaeontological Research, 1984) Olsen, Paul E.; Galton, Peter M.
    The Molteno, Elliot, and Clarens formations comprise the continental Stormberg Group of the Karoo Basin of South Africa and Lesotho. The Molteno Formation contains a well preserved macro- and microfloral assemblage but apparently no vertebrates; the Elliot and Clarens formations contain abundant vertebrates but virtually no floral remains. The vertebrate taxa represented by skeletal remains are listed and divided into two assemblages - the lower Stormberg (lower Elliot) and upper Stormberg (upper Elliot and Clarens) assemblages. The abundant, diagnosable footprint taxa are revised and their names reduced to eight genera. These ichnotaxa also fall into two biostratigraphic zones that parallel the skeletal assemblages. Comparison of the faunal assemblages with those of the European type section strongly suggests that the lower Stormberg assemblage is Late Triassic (Carnian- Norian) in age while the upper Stormberg assemblage is Early Jurassic (Hettangian-Pliens- bachian) in age. Comparisons with other continental assemblages from other areas suggest that the upper Stormberg (upper Elliot and Clarens formations) assemblage broadly correlates with the upper Newark Supergroup of eastern North America, the Glen Canyon of the southwestern United States, and the lower Lufeng Series of China- all thought to be of Early Jurassic age on the basis of floral and/or radiometric evidence. Based on these correlations, previously published paleobiogeographic maps are revised; these show a shift from Late Triassic floral and faunal provinciality to Early Jurassic homogeneity. This shift was synchronous with a widening of the equatorial arid zone.