Wits Evolutionary Studies Institute (ESI)

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Author: search.filters.author.Abdala, FernandoSubject: search.filters.subject.Karoo Basin

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    Cranial morphology and phylogenetic analysis of Cynosaurus suppostus (Therapsida, Cynodontia) from the upper Permian of the Karoo Basin, South Africa
    (Evolutionary Studies Institute, 2018-03) van den Brandt, Marc; Abdala, Fernando
    Non-mammaliaform cynodonts are an important fossil lineage which include the ancestors of mammals and which illustrate the gradual evolution of mammalian characteristics. The earliest cynodonts (‘basal cynodonts’) are known from the late Permian. Cynosaurus suppostus is the second most abundant basal cynodont from the late Permian of the Karoo Basin of SouthAfrica, but is poorly studied, with the most recent description of this taxon being 50 years old. Since then, several important new specimens of this species have been collected, meriting a thorough description of its cranial anatomy and exploration of its interspecific variation. Here we present a comprehensive description of the cranial morphology of Cynosaurus suppostus, producing an updated diagnosis for the species and comparisons amongst basal cynodonts. Cynosaurus is identified by three autapomorphies amongst basal cynodonts: a subvertical mentum on the anterior lower jaw; a robust mandible with a relatively high horizontal ramus; and the broadest snout across the canine region, representing up to 31.74% of basal skull length. One of the new specimens described here preserves orbital scleral ossicles, structures rarely preserved in non-mammaliaform cynodonts. Cynosaurus is now only the third cynodont in which scleral ossicles have been reported.Anupdated phylogenetic analysis of basal cynodont interrelationships recovered Cynosaurus suppostus as a member of the Galesauridae in only two of 16 most parsimonious trees, providing poor support for its inclusion in that family. The majority of known Cynosaurus specimens were collected in a geographically restricted area approximately 150 kilometres in diameter. Most specimens have been recovered from the latest Permian Daptocephalus Assemblage Zone, with only two specimens known from the older Cistecephalus Assemblage Zone.
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    Cranial morphology and taxonomy of South African Tapinocephalidae (Therapsida: Dinocephalia): the case of Avenantia and Riebeeckosaurus
    (Evolutionary Studies Institute, University of the Witwatersrand, 2013-12-18) Guven, Saniye; Rubidge, Bruce S.; Abdala, Fernando
    Tapinocephalid dinocephalians comprise a large component of Middle Permian herbivorous tetrapod biodiversity and are significantly abundant in the Karoo rocks of SouthAfrica. In order to understand the effect of the alleged Middle Permian extinction in the terrestrial record of amniotes a clear understanding of the taxonomic diversity of this group is essential. The tapinocephalids Avenantia kruisvleiensis and Riebeeckosaurus longirostris have in the past been distinguished by a set of characters, including differences in the snout length which was previously interpreted as a key diagnostic character. Further preparation of the material attributed to these species shows that the purported distinguishing characters, such as snout length, low intertemporal crest, and surface thickening of the parietal represent postmortem deformation (including distortion and weathering) or can be interpreted as individual or ontogenetic variation. Accordingly A. kruisvleiensis is synonimized with R. longirostris.