Faculty of Science
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Item The Victoria west: Earliest prepared core technology in the acheulean at canteen kopje and implications for the cognitive evolution of early hominids(Royal Society, 2017-06) Li, H.; Kuman, K.; Lotter, M.G.; Leader, G.M.; Gibbon, R.J.Prepared core technology illustrates in-depth planning and the presence of a mental template during the core reduction process. This technology is, therefore, a significant indicator in studying the evolution of abstract thought and the cognitive abilities of hominids. Here, we report on Victoria West cores excavated from the Canteen Kopje site in central South Africa, with a preliminary age estimate of approximately 1Ma (million years ago) for these cores. Technological analysis shows that the VictoriaWest cores bear similarities to the ‘Volumetric Concept’ as defined for the Levallois, a popular and widely distributed prepared core technology from at least 200 ka (thousand years ago). Although these similarities are present, several notable differences also occur that make the Victoria West a unique and distinctive prepared core technology; these are: elongated and convergent core shapes, consistent blow directions for flake removal, a predominance of large side-struck flakes, and the use of these flakes to make Acheulean large cutting tools. This innovative core reduction strategy at Canteen Kopje extends the roots of prepared core technology to the latter part of the Early Acheulean and clearly demonstrates an increase in the cognitive abilities and complexities of hominids in this time period.Item A re-examination of the enigmatic Russian tetrapod Phreatophasma aenigmaticum and its evolutionary implications(Copernicus GmbH, 2017-02) Brocklehurst, N.; Fröbisch, J.Phreatophasma aenigmaticum is a mysterious tetrapod from the earliest middle Permian of Russia, represented by a single femur. At various times since its original description it has been considered a therapsid synapsid, a pelycosaurian-grade synapsid from the family Caseidae, and most recently a seymouriamorph amphibian. Using up-to-date knowledge of the postcranial morphology and evolution of early synapsids, the specimen is re-evaluated and subjected to cladistic analysis. Seymouriamorph and therapsid affinities are rejected, and a caseid affinity is supported based on the deep intertrochanteric fossa; the widely spaced distal condyles; the short, robust femoral shaft; and the lack of a longitudinal ridge enclosing the posterior margin of the intertrochanteric fossa. When included in two cladistic matrices, the first a global analysis of basal synapsids and the second devoted to caseids, Phreatophasma is found to occupy a basal position within caseids, retaining plesiomorphic characters such as the lack of compression of the anterior condyle of the femur and the almost identical distal extent of the two condyles. The recognition of Phreatophasma as a basal caseid has great implications for the evolution and biogeography of this family. This is only the second example of a caseid from the palaeo-temperate region of Russia, and it is not closely related to the first (Ennatosaurus tecton), implying at least two distinct dispersal events from the palaeoequatorial to temperate latitudes. It also implies that a number of plesiomorphic characteristics of caseids, including small body size and a relatively long femur, were retained as late as the middle Permian, a time when caseids were otherwise represented by large herbivorous taxa.Item Classification and mapping of the woody vegetation of Gonarezhou National Park, Zimbabwe.(AOSIS OpenJournals Publishing AOSIS (Pty) Ltd., 2016-09) Martini, F.; Cunliffe, R.; de Sanctus, M.; D' Ammando, G.; Attorre, F.; Farcomeni, A.Within the framework of the Great Limpopo Transfrontier Conservation Area (GLTFCA), the purpose of this study was to produce a classification of the woody vegetation of the Gonarezhou National Park, Zimbabwe, and a map of its potential distribution. Cover-abundance data of woody species were collected in 330 georeferenced relevés across the Park. These data were used to produce two matrices: the first one using the cover-abundance values as collected in five height layers and the second one based on merging the layers into a single cover value for each species. Automatic classifications were produced for both matrices to determine the optimal number of vegetation types. The two classification approaches both produced 14 types belonging to three macro-groups: mopane, miombo and alluvial woodlands. The results of the two classifications were compared looking at the constant, dominant and diagnostic species of each type. The classification based on separate layers was considered more effective and retained. A high-resolution map of the potential distribution of vegetation types for the whole study area was produced using Random Forest. In the model, the relationship between bioclimatic and topographic variables, known to be correlated to vegetation types, and the classified relevés was used. Identified vegetation types were compared with those of other national parks within the GLTFCA, and an evaluation of the main threats and pressures was conducted. Conservation implications: Vegetation classification and mapping are useful tools for multiple purposes including: surveying and monitoring plant and animal populations, communities and their habitats, and development of management and conservation strategies. Filling the knowledge gap for the Gonarezhou National Park provides a basis for standardised and homogeneous vegetation classification and mapping for the entire Great Limpopo Transfrontier Conservation Area.Item Cranial bosses of choerosaurus dejageri (therapsida, therocephalia): Earliest evidence of cranial display structures in eutheriodonts.(Public Library of Science, 2016-08) Benoit, J.; Manger, P.R.; Fernandez, V.; Rubidge, B.S.Choerosaurus dejageri, a non-mammalian eutheriodont therapsid from the South African late Permian (∼259 Ma), has conspicuous hemispheric cranial bosses on the maxilla and the mandible. These bosses, the earliest of this nature in a eutheriodont, potentially make C. dejageri a key species for understanding the evolutionary origins of sexually selective behaviours (intraspecific competition, ritualized sexual and intimidation displays) associated with cranial outgrowths at the root of the clade that eventually led to extant mammals. Comparison with the tapinocephalid dinocephalian Moschops capensis, a therapsid in which head butting is strongly supported, shows that the delicate structure of the cranial bosses and the gracile structure of the skull of Choerosaurus would be more suitable for display and low energy combat than vigorous head butting. Thus, despite the fact that Choerosaurus is represented by only one skull (which makes it impossible to address the question of sexual dimorphism), its cranial bosses are better interpreted as structures involved in intraspecific selection, i.e. low-energy fighting or display. Display structures, such as enlarged canines and cranial bosses, are widespread among basal therapsid clades and are also present in the putative basal therapsid Tetraceratops insignis. This suggests that sexual selection may have played a more important role in the distant origin and evolution of mammals earlier than previously thought. Sexual selection may explain the subsequent independent evolution of cranial outgrowths and pachyostosis in different therapsid lineages (Biarmosuchia, Dinocephalia, Gorgonopsia and Dicynodontia).