Browsing by Author "Rubidge, Bruce S"
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Item A new middle Permian burnetiamorph (Therapsida: Biarmosuchia) from the South African Karoo filling a gap in the biarmosuchian record(The Evolutionary Studies Institute, 2024-09) Matlhaga, Fonda; Benoit, Julien; Rubidge, Bruce SBiarmosuchia, the most basal therapsid clade, is represented by relatively few specimens known from Permian deposits in Russia and southern Africa. In both the Guadalupian (middle Permian) and Lopingian (late Permian), biarmosuchians represent less than 1% of the fossil record at the specimen level. Here, we describe a new burnetiamorph biarmosuchian, Impumlophantsi boonstrai, based on a partial skull and associated postcrania from the upper Tapinocephalus Assemblage Zone. It is characterized by the presence of a low nasal crest with a unique morphology among burnetiamorphs. Inclusion of this taxon in an updated phylogenetic analysis of biarmosuchians indicates that this specimen is one of the most basal burnetiamorphs, representing the only record of this grade from the middle Permian.Item The bony labyrinth of late Permian Biarmosuchia: palaeobiology and diversity in non-mammalian Therapsida(2017-07) Benoit, Julien; Manger, Paul R; Fernandez, Vincent; Rubidge, Bruce SBiarmosuchia, as the basalmost group of Therapsida (the stem group of mammals), are important for understanding mammalian origins and evolution. Unlike other therapsid groups, the bony labyrinth of biarmosuchians has not yet been studied, despite insightful clues that bony labyrinth morphology can provide to address palaeobiology and phylogeny of extinct animals. Here, using CT scanning, surface reconstruction and a 3D geometric-morphometric protocol of 60 semi-landmarks on the bony labyrinth of 30 therapsids (including three Mammaliaformes), it is demonstrated that bony labyrinth morphology of biarmosuchians is very distinctive compared to that of other therapsids. Despite the primitive nature of their cranial morphology, biarmosuchians display highly derived traits in the structure of the bony labyrinth. The most noticeable are the presence of a long and slender canal linking the vestibule to the fenestra vestibuli, an enlarged and dorsally expanded anterior canal, and the absence of a secondary common crus (except for one specimen), which sets them apart from other non-mammalian therapsids. These characters provide additional support for the monophyly of Biarmosuchia, the most recently recognized major therapsid subclade. Although implications of the derived morphology of the biarmosuchian bony labyrinth are discussed, definitive interpretations are dependent on the discovery of well-preserved postcranial material. It nevertheless sheds light on a previously overlooked diversity of bony labyrinth morphology in non-mammalian therapsids.Item Charles Kimberlin (Bob) Brain - a tribute(Bernard Price Institute for Palaeontological Research, 2000) Rubidge, Bruce SItem Cranial morphology of Jonkeria truculenta (Titanosuchidae, Therapsida, Dinocephalia) and a taxonomic reassessment of the family(Evolutionary Studies Institute, 2023) Jirah, Sifelani; Rubidge, Bruce S; Abdala, FernandoTitanosuchidae are a group of derived herbivorous long snouted dinocephalians currently only known from the South African Karoo. Taxonomic revision of the Titanosuchids, for the first time accompanied by detailed anatomical descriptions and illustrations of representative cranial material, enabled recognition of only two species out of the nine previously recognized. These are Titanosuchus ferox and Jonkeria truculenta. Jonkeria vanderbyli, Jonkeria ingens, Jonkeria haughtoni, Jonkeria parva, Jonkeria rossouwi and Jonkeria boonstrai are here, synonymized with Jonkeria truculenta. The species Jonkeria koupensis is a nomen dubium only identified as Titanosuchid indet. Cranial characters, which modify during ontogenetic development, were, recognized for Jonkeria truculenta, and for the first time an ontogenetic growth series is, presented for this species.Item Dicynodont (Therapsida) bone histology: phylogenetic and physiological implications(Bernard Price Institute for Palaeontological Research, 1993) Chinsamy, Anusuya; Rubidge, Bruce SThe bone histology of humeri of a number of taxonomically well established and easily definable dicynodont genera is described and compared. The bone of Aulacephalodon, Cistecephalus, Dicynodon, Endothiodon, Lystrosaurus, Kannemeyeria and Oudenodon consists of alternating fibro-lamellar and lamellated bone tissue, while that of Diictodon consists only of fibro-lamellar tissue. The presence of fibro-lamellar bone in all the genera studied, indicates that the bone was deposited rapidly, but the occurrence of lamellated bone tissue suggests that all the genera except Diictodon, also had intermittent periods of slow growth. This is the first time that a comparative study of bone histology of different dicynodont genera has been attempted by using one particular bone element to standardise intergeneric comparisons.Item A new vertebrate biozone at the base of the Beaufort Group, Karoo sequence (South Africa)(Bernard Price Institute for Palaeontological Research, 1990) Rubidge, Bruce SA new vertebrate biozone at the base of the Beaufort Group has been identified. This biozone is the oldest vertebrate biozone of the Beaufort Group and contains fossils of various therapsid genera which are more primitive than previously known related forms from the Karoo. The name Eodicynodon - Tapinocaninus Assemblage Zone is proposed as these two genera of therapsid are the most plentiful from this biozone and have not been found in the rocks of the overlying Dinocephalia Assemblage (Tapinocephalus) Zone.