Byrne, Marcus John2017-02-172017-02-171998http://hdl.handle.net/10539/22060Submitted to the Faculty of Science, University of the Witwatersrand, Johannesburg, in fulfilment of the degree of Doctor of Philosophy Johannesburg, South Africa. April 1998Tottenham (Coleoptera: Staphylinidae) were found to co-occur throughout a large expanse of Africa, apparently coexisting in the same habitat. The biology of the three species was investigated to establish the extent of this co-occurrence, and to explore the mechanisms, if any, by which these three species coexist. [Abbreviated Abstract. Open document to view full version]enThesis