1 Peat Dynamics in the Angolan Highlands Mauro Lourenco 830429 A thesis submitted in fulfilment of the academic requirements for the degree of Doctor of Philosophy School of Geography, Archaeology and Environmental Studies University of the Witwatersrand, Johannesburg March 2023 ii DECLARATION I declare that this thesis is my own, unaided work, except where otherwise acknowledged. It is being submitted for the Degree of Doctor of Philosophy in Science at the University of the Witwatersrand, Johannesburg. It has not been submitted before for examination for any degree at this or any other university. ________________________________ Mauro Lourenco 22 March 2023 at Johannesburg iii ABSTRACT The Angolan Highlands is a war stricken, threatened, and under-studied area. The region is hydrologically and ecologically important and supports extensive tropical peatland deposits. Peatland preservation has been acknowledged to address climate change, is sensitive to drought and fire, and is directly influenced by vegetation and hydrological conditions. However, little research has been conducted in the Angolan Highlands. This study addresses gaps in the literature through four key contributions. The first is a critical review of peat definitions: the implications of disparate definitions are detailed, and a new proposed definition for peatlands in the interest of climate science is provided. The second is the first map of peatland extent in the Angolan Highlands, containing details on the age and growth dynamics. The study presents a conservative estimate of peatland extent that is much larger than previously estimated for Angola and is a crucial first step in facilitating the preservation of this deposit. The third contribution is the first historical assessment of drought and vegetation response in the region. This contains a 40-year drought and 20-year vegetation history, demonstrating that drought occurrence is increasing and there is a strong relationship between precipitation and the peatland vegetation region. The fourth contribution is the first assessment of the contemporary (2001-2020) fire regime of these peatlands, and reveals that among all land cover classes, peatlands burn more frequently and at a higher proportion. Investigation into the peat dynamics of the Angolan Highlands indicate that they have critical importance and are naturally resistant to both droughts and fire. Failure to preserve these deposits will have direct implications on the communities, environment, and surrounding areas. Keywords: Angolan Highlands, Carbon, Drought, Fire, Peat, Remote Sensing. iv LIST OF WORKS This study is organised in chapters, four of which are stand-alone research papers that have been accepted for publication in peer-reviewed journals. These are combined with an introduction, study region, methodology, general discussion, and conclusion chapter to present a PhD thesis by publication. The four published papers include: 1. Lourenco M, Fitchett JM and Woodborne S (2022) Peat definitions: A critical review. Progress in Physical Geography: Earth and Environment, DOI: 10.1177/03091333221118353 Current status: Accepted for publication (21 July 2022), available online. Impact factor for 2021-2022: 4.283, five-year impact factor: 5.023. Project leaders Fitchett JM and Woodborne S (30% contribution) conceptualised the scope of this manuscript. Lourenco M (70% contribution) collected and analysed the data, produced the figure and tables in the text, and led the writing of the manuscript and revisions with input from the project leaders. 2. Lourenco M, Fitchett JM and Woodborne S (2022) Angolan highlands peatlands: Extent, age and growth dynamics. Science of The Total Environment, 810, 152315. DOI: 10.1016/j.scitotenv.2021.152315 Current status: Published. Impact factor for 2021: 10.753. Project leaders Fitchett JM and Woodborne S (35% contribution) conceptualised the scope of this manuscript. Lourenco M (65% contribution) collected and analysed the https://doi.org/10.1177/03091333221118353 https://doi.org/10.1016/j.scitotenv.2021.152315 v data, produced the maps, figures, and tables in the text, and led the writing of the manuscript and revisions with input from the project leaders. 3. Lourenco M, Woodborne S and Fitchett JM (2022) Drought history and vegetation response in the Angolan Highlands. Theoretical and Applied Climatology, 151: 115–131 DOI: 10.1007/s00704-022-04281-4 Current status: Published. Impact factor for 2021: 3.409, five-year impact factor: 3.518. Project leaders Fitchett JM and Woodborne S (30% contribution) conceptualised the scope of this manuscript. Lourenco M (70% contribution) collected and analysed the data, produced the maps, figures, and tables in the text, and led the writing of the manuscript and revisions with input from the project leaders. 4. Lourenco M, Woodborne S and Fitchett JM (2023) Fire regime of peatlands in the Angolan Highlands. Environmental Monitoring and Assessment, 195(1): 1–17 DOI: 10.1007/s10661-022-10704-6 Current status: Published. Impact factor for 2021: 3.307, five-year impact factor: 3.420. Project leaders Fitchett JM and Woodborne S (25% contribution) conceptualised the scope of this manuscript. Lourenco M (75% contribution) collected and analysed the data, produced the maps, figures, and tables in the text, and led the writing of the manuscript and revisions with input from the project leaders. https://doi.org/10.1007/s00704-022-04281-4 https://doi.org/10.1007/s10661-022-10704-6 vi Conference presentations: 1. Lourenco M, Fitchett JM and Woodborne S (2021). Towards a peatland Inventory for the Angolan Highlands using Google Earth Engine. Society of South African Geographers and South African Association of Geomorphologists Joint Conference, 6-8 September 2021. 2. Lourenco M, Woodborne S, and Fitchett JM (2022). Drought history and vegetation response in the Angolan Highlands. Society of South African Geographers Biennial Conference, 12-14 September 2022. vii ACKNOWLEDGEMENTS First, I thank my two supervisors, Prof. Jennifer Fitchett, and Prof. Stephan Woodborne for their individual and collective support, guidance, and technical advice throughout my PhD. Through your supervision, you have helped me produce a thesis containing four journal articles accepted for publication in international journals. I am immensely proud of each of these contributions. To Prof. Jennifer Fitchett, thank you for supporting my enthusiasm and curiosity when working in Google Earth Engine. Thank you for your patience and the time taken to help develop my interests into measurable and meaningful outputs. You helped me become a better writer, academic and scientist. Thank you for funding my conference presentations and pushing me to present my work to my fellow colleagues within the school. You taught me to celebrate my academic achievements, to be more organised and productive, and you encouraged me to enter new and exciting avenues of research. Thank you for the countless messages of support, reassurance, and guidance I received throughout. You are a fantastic supervisor, and I am incredibly thankful for your supervision. To Prof. Stephan Woodborne, thank you for helping me to focus on the bigger picture. You helped me develop my critical thinking skills. You taught me to write concisely and communicate my research effectively and with confidence. Thank you for the time you took to create workshop spaces that included skills development, new techniques, theories, and discussions of important science. Thank you for your asking the difficult questions and challenging me to investigate further. Thank you for your insight, the long phone call chats, and advice. You are an excellent supervisor, and I am incredibly thankful for your supervision. viii I thank the anonymous reviewers for their valuable comments and inputs during the review process of each of the four accepted papers. I gratefully acknowledge the support of the National Geographic Okavango Wilderness Project, including the financial support and invitation to attend the Lungui Bungu River expedition in June 2022. I thank Dr. Rainer von Brandis for support, discussions, and feedback during this PhD. I thank Dr. Elhadi Adam for his help and guidance using Google Earth Engine in the early stages of my PhD. I thank Prof. Paida Mhangara and Prof. Diane Grayson for each offering me separate sessional lecturing opportunities at Wits University during my PhD. I thank Wits University for awarding me with the Post Graduate Merit Award and funding my tuition fees over the course of my PhD. I thank my colleagues in the School of Geography, Archaeology and Environmental Studies for your support, encouragement, and insight. I thank my family (my mom, dad, and brother), for supporting me and providing me the opportunity to pursue my PhD. Thank you for your interest in my research and for your unwavering belief in my capabilities. Thank you to my mom for your encouragement and for reminding me that I need to put my education first. Thank you to my dad for reminding me that a PhD does eventually come to an end, and that I need to stay positive. Thank you to my big brother Marcio for your friendship, guidance, insight, and advice. To my Godfather Ricky and his wife Nadia, thank you for your support during my PhD. ix To my extended family and friends that encouraged and celebrated with me throughout my PhD, thank you. I would like to offer special thanks to my grandfather, Vovo Lourenco, who is no longer with us, you will forever have a special place in my heart. Lastly, I would like to offer special thanks to my cousin, Hugo Infante, who although is no longer with us, continues to inspire by the example, dedication and passion he left behind. x TABLE OF CONTENTS Declaration...................................................................................................................ii Abstract.......................................................................................................................iii List of works.................................................................................................................iv Acknowledgements....................................................................................................vii Table of Contents.........................................................................................................x List of Figures.............................................................................................................xiii List of Tables............................................................................................................xviii List of Acronyms and Scientific Terms........................................................................xx Chapter 1: Introduction............................................................................................1 1.1 Background…................................................................................................1 1.1.1 The Angolan Highlands……………………..…..……...................…..…...1 1.1.2 Peat……………………………………………………………………...……2 1.1.3 Remote Sensing……………………………………………...………...…...4 1.2 Rationale……………………………………………...…....................................5 1.3 Study Aim and Objectives..............................................................................6 1.4 Structure of the Thesis...................................................................................7 Chapter 2: Study Region..........................................................................................9 2.1 Introduction....................................................................................................9 2.2 Topography and Drainage...........................................................................11 2.3 Soils and Vegetation....................................................................................13 2.4 Contemporary Climate and Weather..…………...........................................19 2.5 Conclusion...................................................................................................25 Chapter 3: Methodology…………..........................................................................26 xi 3.1 Introduction…..............................................................................................26 3.2 Remote Sensing………………….…............................................................26 3.3 Google Earth Engine……………..................................................................27 3.4 NGOWP Lungui Bungu River expedition…..................................................29 Chapter 4: Peat Definitions: A critical review..........................................................31 4.1 Brief synopsis..............................................................................................31 Chapter 5: Angolan highland peatlands: Extent, age and growth dynamics…........48 5.1 Brief synopsis..............................................................................................48 Chapter 6: Drought history and vegetation response in the Angolan Highlands…..65 6.1 Brief synopsis..............................................................................................65 Chapter 7: Fire regime of peatlands in the Angolan Highlands...............................84 7.1 Brief synopsis……………….........................................................................84 Chapter 8: General Discussion….........................................................................103 8.1 Introduction................................................................................................103 8.2 Peat Preservation in the Angolan Highlands…...........................................103 8.3 Environmental Change in the Angolan Highlands......................................108 8.4 Concerns for the Angolan Highlands region…………….............................111 8.5 Research Limitations…………..……..…....................................................114 8.5.1 Data and analytical limitations for each journal paper chapter……..…115 Chapter 9: Conclusion…......................................................................................118 9.1 Synthesis……............................................................................................118 9.2 Achievement of study aim and objectives…...............................................119 9.3 Future research directions………………………...…..................................122 Comprehensive Reference List................................................................................126 Appendix 1. Chapter 4 Supplementary Material.........................................…………171 xii Appendix 2. Chapter 5 Supplementary Material....……………………………………182 Appendix 3. Chapter 6 Supplementary Material..……………………...………………192 Appendix 4. Chapter 7 Supplementary Material…..…………………………………...198 xiii LIST OF FIGURES Figure 2.1. (a) Angola and its 18 provinces, and (b) the WWF ecoregions occurring in Angola (Olson et al., 2001).........................................................................................10 Figure 2.2. (a) Elevation of Angola depicting the extent of the study site, and (b) elevation of the study site including the WWF HydroSHEDS Basins Level 04 (Lehner and Grill, 2013) and major rivers…………………………………………………………..11 Figure 2.3. Soil map of Angola (from Jones et al. 2013)…..…………………………….14 Figure 2.4. (a) The Cuito Source Lake, Moxico Province. Moist miombo woodland grows on the hillsides adjacent to the lake. Peatland surrounds the source lake, and a narrow band of grassland grows between the peatland and the miombo (From Goyder et al., 2018). Photograph D. Goyder. (b) Lungui Bungu River Source, Moxico Province. Small pool of acidic water filtering out of the surrounding peatland, miombo woodland surrounds the bowl-shaped peatland, and evidence of a recent fire in the background. Photograph R. von Brandis. (c) Lungui Bungu River, Moxico Province. The seep line indicated by a narrow band of white sand between the miombo and the floodplain environment, evidence of small-scale farming within the floodplain environment towards the south of the drone photograph. Photograph J. Guyten............................16 Figure 2.5. (a) Cut section of a peat soil sample extracted by M. Lourenco with a Russian corer. Photograph J. Guyton. (b) Burned (surface), cut, and drained peatland patch, cassava and lavender growing in the background near a small village along the Lungui Bungu River. Photograph M. Lourenco. (c) Drone photograph of a fire event along the Lungui Bungu River floodplain. Photograph J. Guyton................................17 Figure 2.6. Mean annual rainfall in Angola (from Cain, 2017)…..................................21 xiv Figure 2.7. Köppen-Geiger climate classification map for Angola (1980-2016: from Beck et al., 2018)........................................................................................................22 Figure 2.8. (a) Average precipitation (1981-2020) and (b) Average day time land surface temperature (2000-2020) for the delineated study area.................................24 Figure 4.1. (Appearing in the paper as Figure 1): Development of peat definitions through time…………………………………………………………………………………35 Figure 5.1. (Appearing in the paper as Fig. 1): Study site map, (a) Map extent within Angola, (b) Hillshade view of the four riparian Lungui Bungu River cores; samples 1, 2 and 3 lie within the current river floodplain, terrace 1, and sample 4 lies on the relict floodplain, terrace 2, and (c) Map extent for this study, showing the three remaining peat core locations and the hillshade view extent……………………………………….52 Figure 5.2. (Appearing in the paper as Fig. 2): Copernicus Land cover class area and coverage of the Angolan Highlands………………………………………………………53 Figure 5.3. (Appearing in the paper as Fig.3): (a) Landsat 8 and (b) Sentinel 2 RF classifications including coverage and area of each class……………………………...56 Figure 5.4. (Appearing in the paper as Fig. 4): (a) Overlap and non-overlap map showing the extent of panel b and c, (b) and (c) are zoomed in sections of the mapped area………………………………………………………………………………………….57 Figure 5.5. (Appearing in the paper as Fig. 5): Optical, vegetation, standing water occurrence and topographic data of the mapped area from Landsat 8 (a–c), NASA SRTM (d–f) and Sentinel-2 (g–i) sensors……………………………………………...…58 Figure 5.6. (Appearing in the paper as Fig. 6): Distribution plots for L8 and S2 peatland with respect to NDVI, NDWI and SRTM data (elevation and slope). The peaks of each xv distribution plot show the mode of values for individual peatland pixels. The NDVI and NDWI plots relate to each respective peatland class from each RF classification, the SRTM topographical data relates only to L8 peatland.………………………………….59 Figure 5.7. (Appearing in the paper as Fig. 7): Bacon Age-depth profiles for Angolan Highlands peat cores, panels (a–c) represent the age models for CNV: Cuanavale source lake peat, CS: Cuito source lake peat and CU: Cuando source lake cores, respectively. Panels (d–g) represent the age models for Lungui Bungu River cores 1 to 4, respectively……………………………………………………………………………60 Figure 5.8. (Appearing in the paper as Fig. 8): Cross-section of the peat core sampling site at the Lingui Bungu River……………………………………………………………..61 Figure 6.1. (Appearing in the paper as Fig. 1): Study area map showing the study site extent in Angola and a Landsat 8 (USGS, 2021) red, green blue (RGB) optical image of the study site……………………………………………………………………………..69 Figure 6.2. (Appearing in the paper as Fig. 2): (a) The mean annual precipitation (mm/year) over the study site, (b) digital elevation model at 30 m resolution, and (c) the mean cumulative precipitation (mm/month) over the period 1981–01-01 to 2020– 12-31………………………………………………………………………………………...71 Figure 6.3. (Appearing in the paper as Fig. 3): (a–d) The 3-, 6-, 12-, and 24-month SPI for the period 1981–01-01 to 2020–12-31………………………………………………..72 Figure 6.4. (Appearing in the paper as Fig. 4): Mean (a) EVI and (b) NDVI over the study area for the period 2000–02-18 to 2020–12- 31…………………………………..76 Figure 6.5. (Appearing in the paper as Fig. 5): The CHIRPS mean daily precipitation for each month across the Angolan Highlands study area from 1981–01-01 to 2020– xvi 12-31 and the mean NDVI and EVI for each month for the period 2000–02-18 to 2020– 12-31 across the three vegetation regions………………………………………………76 Figure 6.6. (Appearing in the paper as Fig. 6): (a) The mean NDVI and (b) EVI for each vegetation region per month, the total precipitation per month, and the common drought periods over the period (2000–02-18 to 2020–12-31)…………………………77 Figure 7.1. (Appearing in the paper as Fig. 1): (a) Study area map showing the study site extent in Angola and (b) the LULC of the study area………………………………..88 Figure 7.2. (Appearing in the paper as Fig. 2): Peatland sites 1–5 along the Lungu Bungu River…………………………………………………………………………………91 Figure 7.3. (Appearing in the paper as Fig. 3): Fire frequency per pixel over the period 01/01/2001 to 31/12/2020 at 250 m resolution…………………………………………..92 Figure 7.4. (Appearing in the paper as Fig. 4): Burn area per year for the study site over the period 01/01/2001 to 31/12/2020…………………………………………….....92 Figure 7.5. (Appearing in the paper as Fig. 5): (a) Burn proportion and (b) burn area per year for each LULC class over the period 01/01/2001 to 31/12/2020……...……...93 Figure 7.6. (Appearing in the paper as Fig. 6): Proportion of area having specific fire frequencies for each LULC class………………………………………………………….94 Figure 7.7. (Appearing in the paper as Fig. 7): Box-and-whisker plots of the maximum, minimum, average, mode, and inter-quartile range of the annual burned area for each LULC class and all classes per month from 2001 to 2020………………………………96 Figure 7.8. (Appearing in the paper as Fig. 8): Peatland vegetation (average NDVI per month) and burn history (2001–2020) for sites 1–5 (a–e). The black line indicates the time series of average NDVI for each month, a burn value of 1 (red vertical line) xvii corresponds to the month in which the peatland burnt. In some instances, fires occurred over multiple months in the same year, indicating that the peatland site burned in unique locations at slightly different times in the year. (f) Average NDVI per month for all sites during non-burn years (black line) and burn years (red line)……....97 xviii LIST OF TABLES Table 4.1. (Appearing in the paper as Table 1): Peat nomenclature used in definitions…………………………………………………………………………………...37 Table 4.2. (Appearing in the paper as Table 2): Basis for peatland classification systems……………………………………………………………………………………..38 Table 4.3. (Appearing in the paper as Table 3): Peat defined according to depth from various sources……………………………………………………………………………..39 Table 4.4. (Appearing in the paper as Table 4): Threshold proportion of organic carbon, organic matter and ash content that soils must contain to be considered peat………………………………………………………………………………………….40 Table 5.1. (Appearing in the paper as Table 1): Land area coverage of overlap and nonoverlap between the two RF classifications…………………………………………58 Table 6.1. (Appearing in the paper as Table 1): SPI scores and classification (Svoboda et al., 2012)………………………………………………………………………………….70 Table 6.2. (Appearing in the paper as Table 2): Number of months classified as either moderately, severely, or extremely dry per year according to the respective SPI calculations…………………………………………………………………………………74 Table 6.3. (Appearing in the paper as Table 3): Common drought periods according to each SPI calculation and ENSO years………………………………………………...76 Table 6.4. (Appearing in the paper as Table 4): Correlation matrix between vegetation indices and precipitation…………………………………………………………………...77 Table 7.1. (Appearing in the paper as Table 1): Average fire frequency for each LULC class…………………………………………………………………………………………93 xix Table 7.2. (Appearing in the paper as Table 2): Percentage burn proportion per month for each LULC class over the period 01/01/2001 to 31/12/2020. A colour pallet is used to indicate high or low burn proportion……………………………………………………94 xx LIST OF ACRONYMS AND SCIENTIFIC TERMS ~ cal. yr BP: interpolated, calibrated AMS dates calculated using the BACON model; years before present AMS: Accelerator Mass Spectrometry API: Application Programming Interface ASTM: American Society for Testing and Materials CHIRPS: Climate Hazards Group InfraRed Precipitation with Station data CMI: Crop Moisture Index CNV: Cuanavale Source Lake CO2: Carbon dioxide COP: Conference of Parties CS: Cuito Source Lake CU: Cuando Source Lake DEM: Digital Elevation Model DRC: Democratic Republic of the Congo ENSO: El Niño Southern Oscillation ESA: European Space Agency EVI: Enhanced Vegetation Index FAO: Food and Agriculture Organization xxi FAO UNESCO: Food and Agriculture Organization United Nations Educational Scientific and Cultural Organization GEE: Google Earth Engine GIS: Geographical Information System GHG: Greenhouse gas IPCC: The Intergovernmental Panel on Climate Change IPS: International Peatland society ITA: International Trade Administration ITCZ: The Inter-tropical Convergence Zone IUCN: International Union for Conservation of Nature L8: Landsat 8 LB: Lungui Bungu River LULC: Land use land cover m.asl: meters above sea level MODIS: Moderate Resolution Imaging Spectroradiometer NASA: National Aeronautics and Space Administration NDVI: Normalised Difference Vegetation Index (NDVI) NDWI: Normalised Difference Water Index NGOWP: National Geographic Okavango Wilderness Project NIR: Near Infrared xxii NOAA: National Oceanic and Atmospheric Administration PDSI: Palmer Drought Severity Index RCM: Regional Climate Model RDI: Reconnaissance Drought Index RF: Random Forest RGB: Red, green, blue RS: Remote Sensing/ Remotely Sensed RVAA: Regional Vulnerability Assessment and Analysis Programme S2: Sentinel 2 SASSCAL: Southern African Science Service Centre for Climate Change and Adaptive Land Management SF: Surface reflectance SPI: Standardized Precipitation Index SPEI: Standardized Precipitation Evaporation Index SVM: Support Vector Machine SWIR: Shortwave Infrared SRTM: Shuttle Radar Topography Mission TTT: Temperate Tropical Trough UNESCO: United Nations Educational, Scientific and Cultural Organization UNFCCC: United Nations Framework Convention on Climate Change xxiii USGS: United States Geological survey WHO: World Health Organization WMO: World Meteorological Organisation WWF: World Wildlife Fund 1 CHAPTER 1: INTRODUCTION 1.1 Background 1.1.1 The Angolan Highlands Angola has unique habitats and species and is one of the least recognised biodiversity hotspots in the world (Myers et al., 2000; Huntley et al., 2019). Angola is source of many major rivers in southern Africa and is referred to as the “water tower” of the region (Huntley, 2019). These river systems originate from the interior Bié Plateau of the Angola Highlands and flow into large river catchments such as the Cuanza, Cassai (Congo), Lungui-Bungu (Zambezi), Cuito and Cubango (Okavango: Huntley, 2019). The lack of ecological and environmental data and reporting for the upper catchments of the Angolan Highlands is due to historic conflicts in the country (Huntley et al., 2019). The Angolan War of Independence (1961-1974), the Angolan Civil War (1975- 2002) and resultant extensive minefields have prevented access and any scientific exploration or study in the region for over 50 years (Conradie et al., 2016; Midgley and Engelbrecht, 2019). The Okavango Delta, a United Nations Educational, Scientific and Cultural Organization (UNESCO) World Heritage site (International Union for Conservation of Nature: IUCN, 2022), is dependent on precipitation occurring in the highlands of central Angola, where water flows into the Okavango River from two tributaries: the Cuito River and Cubango River (McCarthy et al., 2000; Gumbricht et al., 2004). Concern of the threats to the upper catchments of the Angolan Highlands, and the 2 potential downstream consequences to the Okavango Delta, resulted in the establishment of the National Geographic Okavango Wilderness Project in 2015 (NGOWP; Quammen, 2017). The aim of the NGOWP is to create a network of newly protected areas to conserve the length of the Okavango Catchment (Quammen, 2017). The NGOWP has funded numerous surveys and led a series of expeditions to document the biodiversity and ecology of the Angolan Highlands (Taylor et al., 2018; Goyder et al., 2018; Skelton, 2019; Barber-James and Ferreira, 2019; Van Wilgen et al., 2022). The findings of these expeditions, surveys and studies are intended to provide a scientific foundation to inform the establishment of a system of protected areas within the Angolan Highlands and for the entire Okavango Catchment (Van Wilgen et al., 2022). 1.1.2 Peat Exploratory surveys conducted by the NGOWP between 2015 and 2018 identified peat deposits in the eastern Angolan Highlands (Conradie et al., 2016; Goyder et al., 2018). Peat is a type of organic-rich soil that consists of partially decomposed organic matter derived from plant material (International Peatland society: IPS, 2021). Peat forms when the ground surface is waterlogged due to the interaction between landform, climate, and vegetation (Lindsay, 2016). The term peatland refers to both the peat soil and the terrestrial wetland ecosystem growing on its surface (Dargie et al., 2017). Peatlands, representing at least one third of the global wetlands (Parish et al., 2008), are important ecosystems for biodiversity conservation, carbon storage, climate regulation, biomass production and human welfare (Erwin, 2009; Minasny et al., 2019). Peatlands provide multiple ecosystem services that support the Sustainable 3 Development Goals (Food and Agriculture Organization: FAO, 2020) including purifying water (Frolking et al., 2011; Evers et al., 2016), reduction of flooding and soil erosion (Harenda et al., 2018), aiding in agricultural production and food security (Page et al., 2011), and supporting biodiversity (Minayeva et al., 2017; Xu et al., 2018). Peatlands are the largest natural terrestrial carbon store (Rieley and Page, 2016). Climate change, land use land cover (LULC) change, peatland drainage and fire because of human activities reduce peatland resilience to drought and fire and are linked to peatland degradation, which releases the stored carbon into the atmosphere (Harenda et al., 2018; Minasny et al., 2019). Efforts to reduce the anthropogenic impact on climate has placed great importance on preserving natural carbon sinks such as peatlands (Friedlingstein et al., 2019). Peatlands are estimated to cover 3% of the global land surface (Page et al., 2011; Xu et al., 2018). Although this is a relatively small area, the role of peatlands in the global carbon cycle is significant (Harenda et al., 2018; Loisel et al., 2021). Globally, peat and peatlands have been recognised as essential to multiple international conventions that protect habitats, biodiversity, carbon sinks and reduce greenhouse gas (GHG) emissions (FAO, 2020). The first Peatland Pavilion took place at the 2021 United Nations Framework Convention on Climate Change (UNFCCC) Conference of Parties (COP) 26 (IUCN, 2021). The UNFCCC includes peatlands (organic soils) in its Kyoto Protocol and the Paris Climate Agreement (IUCN, 2021). The Intergovernmental Panel on Climate Change (IPCC) has produced recommendations on national GHG reporting and accounting from drained, rewetted and burning organic soils (IPCC, 2014). Despite its importance, there is no uniform definition for peat or peatland (IPS, 2021). Peatland conservation is dependent on accurate quantification of peatland 4 extent, status, and carbon stock (Minasny et al., 2019; FAO, 2020). Current estimates of global and regional peatland carbon stock and extent are highly varied (Rieley and Page, 2016; Xu et al., 2018). Digital mapping using field observations combined with remotely sensed (RS) imagery and statistical models have been demonstrated to map peatlands more accurately and decrease this uncertainty (Minasny et al., 2019). 1.1.3 Remote Sensing RS products and Geographical Information Systems (GIS) tools and techniques have been successfully utilised for multiple applications including LULC mapping (Mutanga and Kumar, 2019), peatland mapping (Draper et al., 2014; Dargie et al., 2017; DeLancey et al., 2019), vegetation (Huang et al., 2020), drought (AghaKouchak et al., 2021) and fire-related studies (Parks et al., 2018; Bar et al., 2020). The information obtained is used to aid scientists and policy-, and decision-makers (Opolot, 2013), and provides data from inaccessible and extensive areas (Mutanga and Kumar, 2019). The Geo Big Data problem is a view from geospatial and data scientists that require technologies and resources capable of handling large volumes of satellite imagery data (Shelestov et al., 2017). For example, the United States Geological Survey (USGS) has been collecting global earth observation data with frequent intervals since 1972 through the Landsat program (Masek et al., 2020). The data record will continue in future through the joint National Aeronautics and Space Administration (NASA)- USGS Landsat 9 mission which was launched in September 2021 (Gross et al., 2022). Open access to the entire Landsat archive was made available in 2008, however, this archive has been underutilized due to the challenges in collecting, storing, processing, and manipulating this multi-temporal and multi-spectral RS data (Shelestov et al., 5 2017; Teluguntla et al., 2018). In addition, it is impractical to use common image processing software on desktop PC-based systems on data that span large geographic areas over five decades (Teluguntla et al., 2018; DeLancey et al., 2019). Most recently, users have benefitted from the growing availability of high volume, freely accessible RS data and the development of cutting-edge, easy to use machine learning tools that have strong computing capacity (Shelestov et al., 2017; Mahdianpari et al., 2017, 2019; Amani et al., 2020). These advances offer new opportunities for applications at broader spatial and temporal scales, overcoming the limitations of existing methods and products in geospatial sciences (Mutanga and Kumar, 2019; Amani et al., 2020). The establishment of powerful cloud computing infrastructure has been made available through multiple platforms such as NASA Earth Exchange, Amazon’s Web Services, Microsoft’s Azure, and Google cloud platform to address the Geo Big Data problem (Mahdianpari et al., 2019). For instance, Google Earth Engine (GEE), a freely accessible, web-based cloud computing platform that contains numerous geospatial datasets and satellite imagery, allows for algorithm development and processes petabyte-scale data in good time (Hird et al., 2017). 1.2. Rationale The Angolan Highlands are ecologically and hydrologically important (Van Wilgen et al., 2022), supporting peatland deposits (Goyder et al. 2018), and are an essential source water region for the Okavango Delta (Gumbricht et al., 2004) and southern Africa more broadly. Access to the region has been hampered by historical conflicts and persistent minefields, and as a result, it is under-studied (Taylor et al., 2018; 6 Goyder et al., 2018). Peatlands regulate global climate as undisturbed peatlands prevent further climate change (Rieley and Page, 2016). Mapping peatland extent has been documented to facilitate its preservation (FAO, 2020). Estimates of global peatland extent and carbon stock are inconsistent and have high variability (Minasny et al., 2019). This is evident in tropical zones where peatland deposits are comparatively less documented outside of those in Europe and North America (Page et al., 2007; 2011; Rieley and Page, 2016). To address this, the peatland extent in the Angolan Highlands warrants quantification. During periods of drought, peatlands have the potential to become net sources of carbon dioxide as aerobic respiration increases when soil moisture and the water table decrease (Fenner and Freeman, 2011; Lund et al., 2012; Jassey et al., 2018). Having been well documented in the tropics, smouldering combustion of deep peat soil leads to peatland degradation (Page et al., 2002; Page and Hooijer, 2016; Rieley and Page, 2016), causing damage to the environment and human health (Marlier et al., 2013). Undisturbed peatlands are resistant to smouldering combustion due to waterlogging, burning of the surface vegetation does not result in peatland degradation (Vetrita and Cochrane, 2019). Peatland functioning is directly influenced by vegetation and hydrological conditions and is sensitive to drought and fire (Belyea and Malmer, 2004; Page et al., 2011). To address this, an assessment of both drought and the vegetation response to drought, peatland fire regimes and peatland response to fire necessitates investigation. The dynamics and of these peatlands have direct implications on the environment and communities of the Angolan Highlands and surrounding areas, and global climate more broadly. 7 1.3. Study Aim and Objectives The primary aim of this research is to investigate dynamics relating to peatland extent, age and growth, and peatland response to drought and fire in the Angolan Highlands. To address the primary aim of this research, specific objectives of this research are: 1. To determine and map the extent of peatlands in the Angolan Highlands through machine learning. 2. To determine the drought history and the vegetation response to drought in the Angolan Highlands region through assessment of historical satellite data products. 3. To determine the contemporary fire regime of peatlands and peatland response to fire in the Angolan Highlands using fire and vegetation satellite data. 1.4. Structure of the Thesis This thesis is divided into 9 chapters. Chapter 1: Introduction, provides the background, the rationale, and the study aim and objectives. Chapter 2: Study Region, provides a broad synopsis of the Angolan Highlands study region, including descriptions, maps and photographs of the topography, drainage characteristics, soils, vegetation, and contemporary climate. The NGOWP Lungui Bungu River transect took place in June 2022, none of the data collected are reported here, however, photographs obtained during the expedition are presented in Chapter 2. Chapter 3: Methodology, includes details regarding the use of GEE for RS data acquisition, and provides an overview of the NGOWP Lungui Bungu River transect. Chapters 4-7 represent each of the accepted journal papers. Chapter 8: General Discussion, explores three separate themes that cover the four research papers with reference to 8 academic literature, and includes a discussion of the main limitations of the research. Chapter 9: Conclusion, reflects on the extent to which the research aim and objectives have been achieved, the key findings and their significance, and possible avenues for future work. A comprehensive reference list follows the concluding chapter and contains all references from the journal papers and from the thesis chapters. In the case of chapters 4-7, a reference list is included according to the reference style of the respective journal the manuscript was submitted to. The supplementary files from each journal paper appear in the appendices at the end of the thesis rather than at the end of each journal paper. 9 CHAPTER 2: STUDY REGION 2.1 Introduction In this chapter, a description of the World Wildlife Fund (WWF) ecoregions in Angola is presented, followed by the geographic location, topographical and drainage characteristics, soils and vegetation, and the contemporary climate of the study region. Angola is located on the west-coast of south-central Africa, and the country spans 4°22’-18°02’ S, 11°41’-24° 05’ E (Huntley et al., 2019). Angola borders the Democratic Republic of the Congo (DRC) to the north, Zambia to the east, Namibia to the south, and the Atlantic Ocean to the west (Figure 2.1a). Angola covers a total surface area of 1,246,700 km2 (Huntley et al., 2019), and has a population of 33,933,000 (2021), with a population density of 27.22 people per km2 (World Bank, 2022). The WWF terrestrial ecoregions is a global data product that reflects the zonation of the Earth’s terrestrial biodiversity, defining ecological units that share similar environmental conditions, species, and dynamics (Olson et al., 2001). The WWF terrestrial ecoregions showcase the diverse biogeographic and ecological conditions across the country (Figure 2.1b). Dominant ecoregions include the moist forest-savanna mosaics in the north, in the west a 1,600 km coastline contains scarp savanna and woodlands, and desert environments (Olson et al., 2001; Catarino et al., 2020). The miombo woodland ecoregion dominates the central and eastern parts of the country and the southern regions contain arid grasslands, savannas and woodlands along its 1,200 km border with Namibia (Olson et al., 2001; Catarino et al., 2020). 10 Figure 2.1. (a) Angola and its 18 provinces, and (b) the WWF ecoregions occurring in Angola (data from Olson et al., 2001). 11 2.2. Topography and drainage Angola has a heterogeneous topography (Figure 2.2a). In the west, coastal lowlands below 200 m.asl occupy a band between 10–150 km in width and covers 5% of the country (Huntley, 2019). Adjacent to these lowlands, a mountainous escarpment rises to 1000 m.asl and covers 23% of Angola. The plateau is 1000-1500 m.asl and covers 65%, and areas above 1500 m.asl cover 7% of the country (Huntley, 2019). The study area is in the southeast of Angola (Figure 2.2b). This area covers an estimated 61,590 km2 and spans 11°54′-13°54′ S, 18°05′-20°34′ E. The area has an elevation range of 1,117 m.asl in the southeast to 1,678 m.asl in the west. Figure 2.2. (a) Elevation of Angola depicting the extent of the study site, and (b) elevation of the study site including the WWF HydroSHEDS Basins Level 04 (data from Lehner and Grill, 2013) and major rivers. 12 The study area is situated within the Zambezi-Cubango Peneplain which is a vast generally flat area with rivers that meander down a gently dipping plateau (Huntley, 2019). The rivers originate from the northwest watershed with the Cuanza and Congo Basins and flow towards the southeast, eventually reaching the Namibian and Zambian borders (Figure 2.2). The study area forms part of the NGOWP core study region which includes the upper sections of both the Okavango and Zambezi watershed that originate in the Angolan Highlands. The NGOWP extracted peat cores at the Cuito, Cuanavale and Cuando source lakes and near the Lungui Bungu River source. The delineated study area is constrained to the mapped region of the journal paper presented in Chapter 5. This journal paper is a classification map of peatland extent based on the site locations of the peatland cores obtained during the NGOWP expeditions. For consistency, the same study area delineation was used in the journal papers presented in Chapter 6 and 7. The drainage network forms part of the Angolan ‘water tower’, located in the central part of the country (Huntley, 2019). This water tower has nine river basins, of which seven are transnational (Huntley, 2019). The delineated study site includes six major river catchment boundaries: the Lungui Bungu which is a tributary of the Zambezi River and flows east into Zambia; the Cuando which flows south into Namibia, Botswana, Zambia, and Zimbabwe; the Cuito and Cubango which are part of the greater Okavango Catchment flowing southeast into Namibia and Botswana; the Cuanza which flows northwest covering large parts of Angola and the Congo which flows north into DRC. These rivers drain extensive and deep Kalahari sands, due to the filtration caused by the action of the sand layer, the rivers have high clarity and low nutrient levels (Huntley, 2019). 13 2.3 Soils and Vegetation Over 75% of Angola is covered by two main soil groups, the sandy arenosols in the east and the ferralsols in the west and central plateaus of the country (Jones et al. 2013: Figure 2.3). The arenosols are the dominant soil group, covering more than half of Angola (Jones et al. 2013) and most of the study area. The arenosols form part of the Kalahari Basin, the largest body of aeolian sand on earth, extending for 2,500 km from the Cape to the Congo with a maximum width of 1,500 km (Garzanti et al., 2022). These sands cover almost half of Angola, hiding the underlying geology (Huntley, 2019). The quartz grains have no mineral nutrients, little accumulated organic matter, are infertile and have low water-holding capacity (Vainer et al., 2022). The Miombo woodlands ecoregion is particularly well adapted to the arenosols and ferralsols (Olson et al., 2001), and cover most of Angola (Huntley, 2019). The study area also contains alluvial fluvisols within drainage lines, these have high water-retaining capacity and organic content and are suitable for cultivation when not waterlogged (Jones et al. 2013; Huntley, 2019). In addition, gleysol clays are present and typically acidic, waterlogged and occasionally extensive within floodplains (Jones et al. 2013; Huntley, 2019). 14 Figure 2.3. Soil map of Angola (from Jones et al. 2013). Peat soil and peatland deposits were identified during exploratory surveys conducted by the NGOWP (Conradie et al., 2016; Goyder et al. 2018). Tropical peatland deposits have been defined as peatlands which are located between the Tropics of Cancer and Capricorn, including lowland and upland peatlands (Page et al., 2007, 2011). Tropical peatlands occur in Southeast Asia, the Caribbean and Central America, South America, and Africa (Rieley and Page, 2016). Africa has a diversity of peatland depositional environments that vary between sites, the majority of which are minerotrophic (groundwater fed), reflecting the dry climate of the continent (Grundling and Grootjans, 2016). Large ombrotrophic (rainwater fed) bogs exist in wet equatorial 15 regions such as the Congo Basin (Dargie et al., 2017; Davenport et al., 2020). The Angolan Highlands peatlands have formed in upland valleys. Similar peatland formation in upland valleys has also occurred in Rwanda (2,100 m.asl; Hategekimana and Twarabamenya, 2007), Lesotho (2400 m.asl; Trettin et al., 2008), and Burundi (1500 m.asl; Pajunen, 1996). In these high-altitude regions, the conditions for peat formation are comparable to temperate regions (Andriesse, 1988; Page et al., 2011). The Angolan Highlands peatlands have diverse depositional environments, forming in lake margins (Figure 2.4a: Goyder et al., 2018), river floodplains (Figure 2.4b) and river terraces. These peatlands are minerotrophic; with evidence from field observations, drone (Figure 2.4c) and optical imagery, a distinct seep-line indicated by a narrow band of white sand exists parallel to higher ground immediately adjacent to the peatlands. This seep line is the inflow point of groundwater, the valleys are a gentle V-shape, and the peatlands all contour downwards towards the river. 16 Figure 2.4. (a) The Cuito Source Lake, Moxico Province. Moist miombo woodland grows on the hillsides adjacent to the lake. Peatland surrounds the source lake, and a narrow band of grassland grows between the peatland and the miombo (from Goyder et al., 2018). Photograph D. Goyder. (b) Lungui Bungu River Source, Moxico Province. Small pool of acidic water filtering out of the surrounding peatland, miombo woodland surrounds the bowl-shaped peatland, and evidence of a recent fire in the background. Photograph R. von Brandis. (c) Lungui Bungu River, Moxico Province. The seep line indicated by a narrow band of white sand between the miombo and the floodplain environment, evidence of small-scale farming within the floodplain environment towards the south of the drone photograph. Photograph J. Guyten. These peatlands are mostly waterlogged and dark in colour, forming in wet grasslands (Figure 2.5a). The rural communities practice small-scale subsistence agriculture which is reliant on rainfall and environmental conditions (Luetkemeier and Liehr, 2019). Communities often clear the miombo woodlands for cultivation, here the soils (a) (b) (c) 17 are nutrient poor and dry due to the high sand content and high infiltration rates (Hunter and Crespo, 2019; Afonso et al., 2020). In addition, communities cultivate directly on the peatlands (Figure 2.5b), where it is common practice to cut drainage lines and burn the surface vegetation. These subsistence farmers produce low yields of corn, wheat, rice, potatoes, beans, cassava, sugarcane, peanuts, sunflower, sesame, and tobacco (Reyes et al., 2012). The woodlands and grasslands burn frequently (Figure 2.5c). Communities generally burn grasslands in the early dry season (June to July) to aid in hunting practices and clear village surroundings, and woodlands are burnt late in the dry season (August to September) to prepare fields for the growing season (Meller et al., 2022; Teutloff et al., 2022). Figure 2.5. (a) Cut section of a peat soil sample extracted by M. Lourenco with a Russian corer. Photograph J. Guyton. (b) Burned (surface), cut, and drained peatland patch, cassava and lavender growing in the background near a small village along the Lungui Bungu River. Photograph M. Lourenco. (c) Drone photograph of a fire event along the Lungui Bungu River floodplain. Photograph J. Guyton. The Angolan Highlands lie in the Angolan Miombo Woodland ecoregion which contains grasslands, woodlands, savannas and shrublands (Huntley et al., 2019). Barbosa (1970) described the vegetation as dense Zambesian and Congolian miombo woodland with “chanas” or geoxylic-rich grasslands. Within southern Africa, Angola is the least inventoried country for plants, and the east of the country has little to no geo- (a) (b) (c) 18 referenced specimen data (Marshall et al., 2016). Goyder et al. (2018) provide a vegetation checklist and a baseline of plant diversity for the Angolan headwaters, a study that was part of the NGOWP series of expeditions. The species checklist (total of 417) is categorised into four main vegetation types: moist miombo woodlands, swamp forest, seasonally burned savannas, and wetlands (Goyder et al., 2018). Miombo woodlands are tall, closed canopy woodlands dominated by trees in the genera Brachystegia, Julbernardia, and Cryptosepalum (Goyder et al., 2018; Van Wilgen et al., 2022). The miombo is extensive on the hillslopes and dominated by Julbernardia. Miombo growing on the plateau is dominated by Cryptosepalum which can form dense, closed canopy miombo forest rather than miombo woodland. In general, the miombo forests do not have a flammable grass undergrowth that is present in miombo woodland (Goyder et al., 2018). Large scale tree felling of the miombo vegetation has occurred within the Cubango catchment, by contrast, the vegetation in the Cuito catchment remains intact and homogeneous (Mendelsohn, 2019). According to Goyder et al. (2018), swamp forest is rare and only occurs within Moxico Province, unlike the miombo woodlands and the seasonally burned savannas which are extensive. The seasonally burned savannas grow on highly leached Kalahari sand and are described as high-rainfall grasslands (Goyder et al., 2018). The habitat is fire- adapted and is dominated by grasses or geoxylic suffrutices, plants with large underground woody biomass and seasonal above-ground shoots. The factors controlling whether grasses or geoxylic suffrutices dominate are unclear. Maurin et al. (2014) argue fire is an important evolutionary driver, whereas Finckh et al. (2016) 19 provide evidence that frost plays a principal role, with cold air pooling in valley bottoms in the winter dry season and “burning” new shoots. Proximity to the water table limits growth of trees within these savannas (Goyder et al., 2018). Collins et al. (2019) classified these areas as open to sparse woodland and grasslands. Historically, wetlands are under-sampled in Angola (Conradie et al., 2016). According to Goyder et al. (2018), wetlands are typically not botanically diverse, and do not host local endemics. The wide river valleys are characterised by extensive wet grasslands, peatlands, and ox-box lakes (Conradie et al., 2016). These seasonal floodplains along drainage lines are known locally as “chanas” (Van Wilgen et al., 2022) and more commonly referred to as “dambos” across tropical zones of Africa (Midgley and Engelbrecht, 2019; Skelton, 2019; Moore et al., 2022). The impeded drainage and high precipitation in the rainy season cause temporarily waterlogged soils in the valleys that support humid grassland borders with humic topsoil and dwarf shrubs and prevent the development of miombo woodland (Conradie et al., 2016). The river source lakes have peat accumulations at their margins and the river valleys also contain peat deposits (Goyder et al., 2018). Although described as not botanically diverse, the species checklist provided by Goyder et al., (2018) contains a total of 115 plant species identified in the wetland habitat of the Cuito catchment alone, 94 of which are associated with peat soils. 2.4 Contemporary Climate and Weather The climate and weather of Angola is diverse, owing to topographic, atmospheric, and oceanic factors. There is a mean annual temperature difference of 4 °C (24.7 °C near the Equator versus 20.7 °C near the Tropic of Cancer), decreasing from north to south 20 (Huntley, 2019). Temperatures generally decrease with altitude, driven by the adiabatic lapse rate. There are major atmospheric systems over central and southern Africa that influence rainfall patterns across the country (Nicholson, 2018). The Inter- tropical Convergence Zone (ITCZ) is a belt of low pressure which circles the globe near the equator (Schneider et al., 2014). This is where the trade winds from the Southern and Northern Hemispheres converge, generating convective activity which drives moist conditions that are typical of the tropics (Nicholson, 2018). Seasonal shifts in the location of the ITCZ affects the precipitation of many equatorial countries, resulting in distinct wet and dry seasons common in the tropics, rather than cold and warm seasons that occur in higher latitudes (Schneider et al., 2014; Nicholson, 2018). The ITCZ is on the equator during mid-autumn and mid-spring and migrates towards the Tropic of Cancer in the austral winter. The wet season starts when the ITCZ and the Congo Air Boundary shift over the north of Angola in early summer and reach the south of the country in late summer (Huntley, 2019). Mean annual rainfall is generally highest in the north and northeast (1680 mm/ year) and decreases towards the southwest (100 mm/year), high mean annual rainfall (1500 mm/ year) occurs in the high-altitude regions near Huambo Province (Figure 2.6; Cain, 2017). 21 Figure 2.6. Mean annual rainfall in Angola (from Cain, 2017). The climate is seasonal; hot and wet summers occur from October to May and mild to cool and dry winters occur from June to September (Huntley, 2019). The Köppen- Geiger climate classification (1980-2016) illustrates distinct climate zones in Angola (Figure 2.7; Beck et al., 2018). Tropical, savanna (Aw) covers large expanses of the central and northern parts of the country. The coastline is mostly arid, containing hot deserts (BWh) in the southwest (Beck et al., 2018). The Angola and Benguela Currents have a stabilising effect on the lower atmosphere and prevent cloud formation, resulting in the evolution of the Namib Desert (Lima et al., 2019). There are 22 two high-pressure systems that shift along with the ITCZ over the Atlantic Ocean and southern Africa: The Botswana Anticyclone and the South Atlantic Anticyclone. In winter, the anticyclones block the southward movement of moist air and prevent cloud formation. In summer, the anticyclones shift south and rainfall returns (Huntley, 2019). During the months spanning winter through to early summer, strong east-west winds are induced by the Botswana Anticyclone, picking up dust from the arid steppe and contributing to the Namib dunes (Huntley, 2019). The study area is characterized by Tropical, savannah (Aw), Temperate, dry winter, hot summer (Cwa) and Temperate, dry winter, warm summer (Cwb) climates which span the plateau (Beck et al., 2018). Figure 2.7. Köppen-Geiger climate classification map for Angola (1980-2016: adapted from Beck et al., 2018). Studies that focus on Angola’s climate have been hampered due to the collapse of an extensive network of metrological stations (Huntley, 2019). Between 1974 and 2010, Angola’s climate network was reduced from 225 climatological posts (Silveira, 1967) 23 to zero, and 29 synoptic stations to 23 stations (Government of Angola, 2013). This is demonstrated in the study by Silveira (1967), a critical assessment of Angola’s climate, that contained data from 184 stations across all 18 provinces. In addition, the study by Azevedo (1972), to map and classify the climatic regions, was based on this considerable national dataset, and still provides an accurate representation of Angolan bio-climatic systems (Huntley, 2019). In the case of Peel et al. (2007), a more recent study which provides an updated world map of the Köppen-Geiger climate classification, only five temperature and sixteen precipitation stations were used for Angola. Furthermore, records of extreme minimum temperatures and frost in Angola are absent (Zigelski et al. 2019). These factors, in combination with fires and herbivory, are important to the floristic diversity and composition of the country (Zigelski et al. 2019). Due to the sporadic national meteorological station coverage and lack of reliable climatic data, this study makes use of climatic data obtained from RS products (Figure 2.8). Historical climate data from weather stations, specific to the study area, are unavailable. Distinct differences in precipitation and land surface temperatures occur within the study area. Historical precipitation and land surface temperature data for the region were collated from the GEE platform. Precipitation data were obtained from the Climate Hazards Group InfraRed Precipitation with Station data (CHIRPS) 30+ year quasi-global rainfall dataset (Funk et al., 2015). CHIRPS synthesises 0.05° resolution satellite imagery with in-situ station data to create a gridded rainfall time series (Funk et al., 2015). Land surface temperature data were obtained from the Terra Moderate Resolution Imaging Spectroradiometer (MODIS) Land Surface Temperature/ Emissivity 8-Day (MOD11A2) Version 6.1 product (Wan et al., 2021). This product 24 provides an average 8-day per-pixel land surface temperature and emissivity with a 1 km spatial resolution in a 1,200 by 1,200 km grid (Wan et al., 2021). Figure 2.8. (a) Average precipitation (1981-2020) and (b) Average day time land surface temperature (2000-2020) for the delineated study area. The mean annual precipitation over the study area is 1113 mm for the period 1981- 01-01 to 2020-12-31. The highest annual precipitation (1295 mm) is recorded in the northwest and northeast regions of the study area (Figure 2.8a). Precipitation in the region is closely related to latitude and altitude. Both precipitation and altitude decrease towards the southeast where the mean annual precipitation is lowest (904 mm). The landscape has distinct topographical features dominated by river processes; high elevations exist adjacent to low lying river valleys. Land surface temperatures reflect these topographical features and are closely related to vegetation (Figure 2.8b). On average, miombo woodlands which grow on high elevations have cooler (24.5 °C) 25 land surface temperatures in comparison to the warmer valleys at lower elevations (33.6 °C). 2.5 Conclusion Angola is a large country on the western coast of southern Africa. The important components of Angola’s diverse ecoregions, topographical and drainage characteristics, soils, vegetation, and contemporary climate are detailed. These range from the Namib desert in the southwest, to the arid savannas of the coastal and southern areas, to the steep Angolan escarpment. Above the escarpment, high mountains contain Afromontane grasslands and forests. These mountainous areas are the source of several large rivers which drain into multiple neighbouring countries. Extensive miombo woodlands dominate the plateau and peneplains. Mean annual rainfall varies from below 100 mm in the southwest to over 1600 mm in the north of Angola. The precipitation and temperature across the country are closely linked to global and regional synoptic systems creating distinct wet and dry seasons. In addition, the topography and ocean currents have an influence on climate. Within the study area, the landscape contains distinct vegetation types, dominated by miombo woodlands. The peatlands which grow in the river valleys are controlled by multiple factors including groundwater, climate, topography, and vegetation. These peatlands provide extremely important ecosystem services as they purify water, limit river flow, provide fertile soil for agriculture, are a fuel source, and store carbon. Although the study area is rural, the strong anthropogenic influence on fire, wetland drainage and tree clearing are a threat to this region. This underscores the importance of investigating the peat dynamics within this area of Angola. 26 CHAPTER 3: METHODOLOGY 3.1 Introduction This chapter provides a brief methodological background to the use of Google Earth Engine in remote sensing studies. In addition, an overview of the NGOWP Lungui Bungu River transect conducted in June 2022 is included. The specific analytical methods used in the analysis of peat extent, of drought and fire dynamics are detailed in each of the respective journal papers. 3.2 Remote Sensing The Angolan Highlands are an extensive, understudied area that is largely inaccessible because of landmine presence following historic conflicts (Taylor et al., 2018; Goyder et al., 2018). Since the inception of the NGOWP in 2015, although conducting the most extensive scientific exploration across the region for 50 years (Conradie et al., 2016; Midgley and Engelbrecht, 2019), access to the region remains difficult and costly. RS has several advantages over traditional mapping and data collection approaches, field measurement techniques are limited by accessibility, scale, and cost, whereas remote sensing is quicker, more cost effective, and provides detailed information at a regional scale (Lees et al., 2018; Wu et al., 2014). Satellite data cover areas that are inaccessible and the repeat data collection enables effective environmental monitoring of dynamic phenomena such as precipitation, vegetation growth and fire (Mutanga and Kumar, 2019; Shiklomanov et al., 2019). 27 There are a wide range of RS datasets available that are often used in combination: multi-sensor approaches are highly useful for a variety of environmental monitoring research topics and have been used to identify and discriminate peatland from other wetland features (Hird et al., 2017; Mahdianpari et al., 2019; DeLancey et al., 2019). The availability of big data from earth observation products and the advances in machine learning, has provided further opportunities for new methods to aid in earth environmental monitoring (Gorelick et al., 2017; Amani et al., 2020; Yuan et al., 2020). Over the last decade, substantial progress in earth sciences has been observed, these new advances have outperformed traditional RS models with considerable improvement in performance (Tamiminia et al., 2020; Yuan et al., 2020). 3.3 Google Earth Engine In recent years, following the significant advancement in sensors and increase in the number of RS datasets, multiple challenges to users working with big data have been created (Chi et al., 2016; Tamiminia et al., 2020). The challenges include big data computing, collaboration, and methodologies, as well as the appropriate data identification, deployment, representation, fusion, visualisation, and interpretation (Chi et al., 2016; Gorelick et al., 2017; Amani et al., 2020). The development of a safe, efficient, and advanced cloud computing platform was one of the most important requirements to provide a comprehensive solution to these challenges (Gorelick et al., 2017; Amani et al., 2020). Cloud computing platforms provide infrastructure, storage services, datasets, and software packages that enable the performance of a supercomputer on a standard device (Yuan et al., 2020; Amani et al., 2020). 28 GEE is a cloud computing platform launched by Google in 2010 (Tamiminia et al., 2020). GEE uses Google’s computational infrastructure and has a large collection of open access satellite imagery and RS datasets that are constantly updated (Gorelick et al., 2017; DeLancey et al., 2019; Tamiminia et al., 2020). GEE is freely accessible and the most popular big data processing platform (Amani et al., 2020). Users can access GEE through an internet-based Application Programming Interface (API) and a web-based interactive development environment (Mahdianpari et al., 2019). Furthermore, users do not need to download the large datasets and GEE has an automatic parallel processing and fast computational platform that effectively deals with big data at a petabyte scale (Gorelick et al., 2017; Amani et al., 2020). In addition to its computing and storage capacity, several well-known machine learning algorithms have been implemented, and users are able to develop and share their own algorithms easily (Chi et al., 2016; Tamiminia et al., 2020). The data outputs and map exports are complimentary to existing RS and GIS software packages (Chi et al., 2016; DeLancey et al., 2019). Considering the trends of GEE use in research over the recent past, it is projected that users will more frequently use GEE as the main cloud computing service in future (Tamiminia et al., 2020). In this study, GEE was used as the primary platform for RS data extraction and storage and used in some analytical approaches. Map visualisations and calculations were performed using ArcMap software. Sample GEE scripts were obtained from the dedicated GEE developers guide and API Javascript tutorials. These were revised for the study site and specific date ranges for each journal paper chapter. During this study, an ethics clearance was not needed as the data collected was solely obtained from freely accessible RS datasets on GEE. 29 3.4 NGOWP Lungui Bungu River expedition To ensure long-term, sustainable protection for the Okavango watershed, the NGOWP has been conducting surveys, gathering scientific data concerning the river systems and collaborating with local communities, non-government organisations and the governments of Angola, Namibia, and Botswana (National Geographic Society, 2022). Since 2015, the NGOWP have completed multiple expeditions, performing river transects across the length of the catchment from the source waters in the Angolan Highlands to the Makgadikgadi Pan in Botswana (National Geographic Society, 2022). The NGOWP core study region has since expanded to other major river basins originating from the Angolan Highlands including the Zambezi catchment. The first NGOWP expedition conducted in Angola following the COVID-19 global pandemic took place in June 2022. The transect for this field work as part of the PhD, started at the Lungui Bungu River source, and continued east, along the river, until the Angola- Zambia border. The Lungui Bungu River is a major tributary to the Zambezi River. Prior to this expedition, observations of the landscape were limited to RS satellite datasets as this was the candidates first visit to the study area. In previous NGOWP expeditions, peat cores were collected on the river floodplain near the Lungui Bungu River source. Following Accelerator Mass Spectrometry (AMS) radiocarbon dating, important data regarding peatland formation was obtained. With this background knowledge, a peatland classification map for the region was generated prior to this expedition. The Lungui Bungu River transect provided an opportunity to validate this peatland classification. In addition, observations of the peatland landscape revealed further insight of the threats to these deposits and provided additional verified peatland 30 extent data that was used in Chapter 7. The specific details regarding the data and methodologies used are mentioned in each journal paper chapter. The principal aim of this expedition was to conduct the first known scientific exploration of the length of the Lungui-Bungu River. Selected outcomes and achievements of the expedition are described in this thesis. However, the primary results of this expedition fell outside of the scope of this work and therefore do not appear in this thesis. Importantly, this provided the opportunity to visit the study site and to understand the context in which the RS work was done. Outcomes specific to the peatlands include an additional 17 core samples of peat soil that were collected along the length of the river. The cores were collected using a Russian peat corer and have been radiocarbon dated at the iThemba LABS. In addition, water quality data using a multiparameter probe and water discharge using an acoustic doppler current profiler were collected every 10 km along the river length. Biodiversity counts, evidence of human interaction with the river (for example, fishing nets) and fire events near the river were recorded along the transect. River invertebrate biodiversity assessments were performed every 30 km along the river. 31 CHAPTER 4: PEAT DEFINITIONS: A CRITICAL REVIEW 4.1 Brief synopsis This critical review presents a brief history on peatland definitions through time. The review highlights the current state and discrepancies between definitions concerning peatland nomenclature and classifications. Multiple disparate definitions are presented with specific focus on the criteria concerning minimum depth, organic carbon, organic matter, and ash content. This study presents a new definition for peat, the motivation behind this definition is for conservation purposes and from the perspective of climate science, preservation, and carbon accounting. This is an important contribution as the literature concerning peat and peatland definitions is inconsistent, causing multiple implications, including the quantification of global peatland carbon stock. Author contributions: Contributor role Author contribution Conceptualisation Lourenco M, Fitchett JM, and Woodborne S Methodology Lourenco M, Fitchett JM, and Woodborne S Validation Lourenco M Formal analysis Lourenco M Investigation Lourenco M Resources Lourenco M Data curation Lourenco M Writing – the original draft preparation Lourenco M Writing – review and editing Lourenco M, Fitchett JM, and Woodborne S Visualisation Lourenco M Supervision Fitchett JM and Woodborne S Project leaders Fitchett JM and Woodborne S According to the above-mentioned CRediT system the author contribution was calculated as follows: Lourenco M – 70% 32 Fitchett JM and Woodborne S – 30% Submitted to: Progress in Physical Geography Impact factor for 2021-2022: 4.283. Submission date: 29 March 2022 Revision date: 15 July 2022 Accepted: 21 July 2022 Published online: 4 October 2022 Available from: https://journals.sagepub.com/doi/full/10.1177/03091333221118353 The version to follow is exactly as published in the journal on 4 October 2022, with the addition of contiguous page numbers for this thesis. https://journals.sagepub.com/doi/full/10.1177/03091333221118353 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 CHAPTER 5: ANGOLAN HIGHLANDS PEATLANDS: EXTENT, AGE AND GROWTH DYNAMICS 5.1 Brief synopsis This study presents the first classification of the Angolan Highlands peatlands, including a map of the peat extent. In addition, analysis of the age and growth dynamics of these deposits are presented. This is a multi-disciplinary study that incorporates RS datasets, machine learning and GEE with AMS radiocarbon dates of peat cores. Tropical peatland deposits are poorly mapped and documented within the literature. This is an important contribution as the study provides a first conservative estimate of the extent of these peatlands, contains details of peatland characteristics with respect to topographical data and RS indices concerning vegetation and standing water and suggests possible dates for peatland growth initiation, peatland growth dynamics, potential carbon storage, and threats to this deposit. Author contributions: Contributor role Author contribution Conceptualisation Lourenco M, Woodborne S, and Fitchett JM Methodology Lourenco M, Fitchett JM, and Woodborne S Validation Lourenco M Formal analysis Lourenco M, Fitchett JM, and Woodborne S Investigation Lourenco M Resources Lourenco M Data curation Lourenco M Writing – the original draft preparation Lourenco M Writing – review and editing Lourenco M, Fitchett JM, and Woodborne S Visualisation Lourenco M Supervision Fitchett JM and Woodborne S Project leaders Woodborne S and Fitchett JM 49 According to the above-mentioned CRediT system the author contribution was calculated as follows: Lourenco M – 65% Fitchett JM and Woodborne S – 35% Submitted to: Science of the Total Environment Impact factor for 2021: 10.753 Submission date: 14 July 2021 Revision dates: 20 October 2021 and 1 December 2021 Accepted: 7 December 2021 Published: 13 December 2021 Available from: https://www.sciencedirect.com/science/article/pii/S0048969721073915 The version to follow is exactly as published in the journal on 13 December 2021, with the addition of contiguous page numbers for this thesis. https://www.sciencedirect.com/science/article/pii/S0048969721073915 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 CHAPTER 6: DROUGHT HISTORY AND VEGETATION RESPONSE IN THE ANGOLAN HIGHLANDS 6.1 Brief synopsis In this study, an investigation of the drought dynamics for the Angolan Highlands are presented from the analysis of a 40-year CHIRPS precipitation and a 20-year MODIS vegetation growth record. The study reveals the contemporary seasonality of precipitation and vegetation growth for the region. It presents the 40-year drought history and the describes the relationship between drought and ENSO and regional synoptic systems. The implications of drought are discussed in this study, with specific reference to peatland vegetation growth under drought conditions and future climate scenarios. This is an important contribution as the study provides the first historical assessment of drought in the Angolan Highlands, a region which contains vulnerable rural populations that are dependent on rain-fed subsistence agriculture. Author contributions: Contributor role Author contribution Conceptualisation Lourenco M, Fitchett JM and Woodborne S Methodology Lourenco M, Fitchett JM, and Woodborne S Validation Lourenco M Formal analysis Lourenco M Investigation Lourenco M Resources Lourenco M Data curation Lourenco M Writing – the original draft preparation Lourenco M Writing – review and editing Lourenco M, Fitchett JM, and Woodborne S Visualisation Lourenco M Supervision Fitchett JM and Woodborne S Project leaders Fitchett JM and Woodborne S 66 According to the above-mentioned CRediT system the author contribution was calculated as follows: Lourenco M – 70% Fitchett JM and Woodborne S – 30% Submitted to: Theoretical and Applied Climatology Impact Factor for 2021: 3.409 Submission date: 28 February 2022 Revision dates: 31 May 2022, 3 September 2022, and 31 October 2022 Accepted: 31 October 2022 Published online: 10 November 2022 Available from: https://link.springer.com/article/10.1007/s00704-022-04281-4 The version to follow is exactly as published in the journal on 10 November 2022, with the addition of contiguous page numbers for this thesis. https://link.springer.com/article/10.1007/s00704-022-04281-4 67 68 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 CHAPTER 7: FIRE REGIME OF PEATLANDS IN THE ANGOLAN HIGHLANDS 7.1 Brief synopsis Using MODIS fire and vegetation data in combination with the LULC map presented in Chapter 5, this study presents the first assessment of the contemporary fire regime of peatlands in the Angolan Highlands. Fire and associated degradation have been extensively documented in tropical peatland deposits in southeast Asia, with little to no reporting for African tropical peatlands. This is an important contribution as the study provides a first assessment of the fire regime of peatlands, including the implications of fire for peatlands, and discusses fire management, future research, and area conservation for the Angolan Highlands. Author contributions: Contributor role Author contribution Conceptualisation Lourenco M, Fitchett JM, and Woodborne S Methodology Lourenco M, Fitchett JM, and Woodborne S Validation Lourenco M Formal analysis Lourenco M Investigation Lourenco M Resources Lourenco M Data curation Lourenco M Writing – the original draft preparation Lourenco M Writing – review and editing Lourenco M, Fitchett JM, and Woodborne S Visualisation Lourenco M Supervision Fitchett JM and Woodborne S Project leaders Fitchett JM and Woodborne S According to the above-mentioned CRediT system the author contribution was calculated as follows: Lourenco M – 75% Fitchett JM and Woodborne S – 25% 85 Submitted to: Environmental Monitoring and Assessment Impact factor for 2021: 3.307 Submission date: 22 August 2022 Revision date: 19 October 2022 Accepted: 25 October 2022 Published: 7 November 2022 Available from: https://link.springer.com/article/10.1007/s10661-022-10704-6 The version to follow is exactly as published in the journal on 13 December 2021, with the addition of contiguous page numbers for this thesis. https://link.springer.com/article/10.1007/s10661-022-10704-6 86 87 88 89 90 91 92 93 94 95 96 97 98 99 100 101 102 103 CHAPTER 8: GENERAL DISCUSSION 8.1 Introduction This chapter presents the general discussion, divided into three themes cross cutting the four research papers (Chapters 4-7), namely peat preservation in the Angolan Highlands, environmental change in the Angolan Highlands, and concerns for the Angolan Highlands region. In addition, this chapter integrates and expands on the findings and connections between each research paper with reference to literature. During the PhD, Chapter 5 was the first to be published, followed by Chapter 4. Chapters 6 and 7 were in review at the same time and published shortly after one another. Although having similar vegetation, precipitation, and fire time-series records, the journal articles that appear in Chapters 6 and 7 were submitted as standalone research papers. A discussion of the limitations of this study is included in this chapter. 8.2 Peat preservation in the Angolan Highlands Peatlands have been identified in at least 175 countries, occurring in polar and tropical regions and every climatic zone (Lindsay, 2010; Bain et al., 2011; IPS, 2020). Due to available resources and widespread peatland coverage, peatlands have been mapped extensively in the northern hemisphere; however, shortfalls exist in adequate peatland distribution maps both for the southern hemisphere and the tropics (Page et al., 2011; Evans et al, 2014). As a result, the dissemination of global peatland distribution maps is biased towards the northern hemisphere (Wu et al., 2017; Xu et al., 2018). Global peatland maps and estimates of peatland extent are also inconsistent as they are produced from several differing data sources at global, regional, and national levels 104 (Wu et al., 2017; Xu et al., 2018). These collection approaches are both dependent and constrained to high quality peatland extent data that are only available for a small selection of countries and regions, including Canada, Sweden, and West Siberia (Wu et al., 2017). The first peatland map of the Angolan Highlands (Chapter 5) provides a conservative estimate (1,634 km2) of peatland extent for the region. The best estimate (2,640 km2) of peatland extent for Angola reported by Page et al. (2011) makes no mention of these deposits, and it is likely that the estimate is an addition to peatland extent reported for Angola. This estimate is a crucial first step in providing the peat carbon inventory and facilitates conservation efforts for the region and surrounding basins. In addition, it provides updated peatland data for Angola that may be used in updated estimates of global peatland extent. Peatlands store more carbon than the above-ground carbon stored in all the world’s forests (Page et al., 2011; Bain et al., 2011). Despite this, peatlands have received less attention under the UNFCCC (Ramsar, 2002; Evans et al, 2014), and have only recently received substantial recognition at the 2021 UNFCCC COP26 (IUCN, 2021). The focus of the literature over the last decade has steered toward peatland preservation (Finlayson and Milton, 2018; Rieley and Page, 2016). This is facilitated by accurate classification of peatland extent (Minasny et al., 2019; FAO, 2020) and quantification of peatland carbon (Law et al., 2015; Xu et al., 2018). In Chapter 4, the study extends this focus, stating that discrepancies in peat and peatland definitions are negatively influencing efforts in quantifying global peat carbon stock. In this study, a new definition for peatland is proposed, motivated by climate science, preservation, 105 and carbon accounting. The implication is that a common definition for peat and peatland will also contribute towards its preservation. Chapter 5 does not include the conservative definition proposed in Chapter 4 as it was written and accepted before the comprehensive review of peat and peatland definitions. There are several anthropogenic practices that threaten the preservation of the Angolan Highlands peatland deposits, including extensive fires, slash and burn agriculture (also known as shifting agriculture), peat fuel extraction, wetland drainage, and overgrazing (Conradie et al., 2016; Taylor et al., 2018). As calculated in Chapter 7, the peatlands burn more frequently, have the smallest proportion of area which does not burn, and have the largest average proportion of burnt area per year among all the LULC classes. Peatland functioning is dependent on continual recycling of organic matter under waterlogged, anaerobic conditions (Lawson et al., 2015). Conditions where either vegetation is removed, or the water table is lowered are major threats to peatland growth and functioning (Rieley and Page 2016). Human interference on peatlands can be highly detrimental (Brevik and Homberg, 2004; Hooijer et al., 2010) as drainage, peat extraction, inappropriate burning, and conversion to agriculture all lower the water table (Bain et al., 2011; Page et al., 2011). The more intensively the peatland is disturbed, the quicker the peat degrades, oxidises and releases GHGs (Bain et al., 2011; Evans et al., 2014; Lawson et al., 2015). Compromised peatlands become a major source of GHG emissions which may continue for many years until all the peat is lost (Joosten 2011; Bain et al., 2011). The combination of extensive area covered by peatlands and potential long-term emissions makes the climate effect of degraded peatlands fundamentally distinct from other 106 ecosystems (Joosten 2011; Bain et al., 2011), requiring consistent monitoring and unique preservation strategies (FAO, 2020). Peatlands are sensitive to temperature, precipitation, and prolonged periods of drought (Belyea and Malmer, 2004). In Chapter 6, drought events in the region are shown to increase through time and negative SPI trends indicate that drought occurrence and frequency are likely to increase in future. The vegetation response to drought was calculated for the study area, and more specifically for the vegetation within the valley environment which contains much of the peatlands classified in Chapter 5. Under drought conditions, both soil moisture and the water table decrease, facilitating the increased potential for aerobic respiration, causing peatlands to become net sources of CO2 (Fenner and Freeman, 2011; Lund et al., 2012; Jassey et al., 2018). The relationship between drought and ecosystem response is often non-linear (Jassey et al., 2018), however, tipping points have been identified for individual peat forming species in response to drought, when reached these thresholds that are detrimental to the ecological functioning of the peatland (Lund et al., 2012; Lamentowicz et al., 2019). Peatlands rarely burn when waterlogged (Rieley and Page, 2016), however, if desiccated due to drought and or drainage, they are more easily combustible (Vetrita and Cochrane, 2019). Peatland loss and degradation due to the combustion of deeper peat has been documented in boreal and tropical peatlands across the globe (Rieley and Page, 2016; Turetsky et al., 2015). The results presented in Chapter 7 suggest that peatland burning within the five intact peatland sites are surface vegetation fires, where the above-ground peat forming vegetation is burnt and recovers shortly after 107 burning. Smouldering (deep peat) fires would likely result in peatland degradation (Vetrita and Cochrane, 2019), where the surface vegetation would not be able to recover post burn and continue to burn year on year. It is critical that these peatlands remain undisturbed and are protected because they have a natural resistance to smouldering combustion and degradation, protecting the deeper peat layers that have accumulated carbon over thousands of years. Within undisturbed peatlands throughout the world, most of the peatland carbon stock is typically protected from smouldering, but drying because of human activity, drought and climate change lowers the water table, exposing the deeper peat to smouldering combustion and degradation (Vetrita and Cochrane, 2019). According to Regional Climate Models (RCMs) for Angola, the Angolan Highlands are projected to become both drier (decrease in precipitation of up to 4%) and warmer (increase in air temperature of up to 4.9 °C) by 2100 (Carvalho et al., 2017). Continued population growth, subsistence agriculture and human pressure on the land are increasing in unprotected rural areas of Angola such as the Angolan Highlands (Catarino et al., 2020). Local communities are likely to target the moist and organic rich peatlands for cultivation in a warmer and drier climate future (Humpenöder et al., 2020), lowering the water table and exposing deeper peat to fire and degradation. Multiple countries have developed national peatland strategies to promote preservation to ensure continued functionality (Andersen et al., 2017). Standard peatland management practices include the promotion of peat forming vegetation, prevention of water loss and water pollution and reduction of peat extraction, cultivation, and drainage (Dohong et al., 2017). Context-driven strategies should be 108 considered with the input of local ecological and indigenous knowledge, communities, and stakeholders (Van Noordwijk et al., 2014; Fleming et al., 2021). 8.3 Environmental change in the Angolan Highlands Over the last decade, unprecedented changes in the human and biophysical environments have occurred (Loisel et al., 2021). These include the increase of fossil fuel emissions and rising global temperatures, which place greater importance on carbon sinks (Friedlingstein et al., 2019). Reducing the atmospheric abundance of carbon dioxide (CO2) and other GHGs is required to mitigate the risks of climate change (Houghton, 2002; Archer et al., 2009). Peatlands contribute towards this mitigation as they are an important natural reservoir of global carbon (Ramsar, 2002). In the 6th Assessment report of the IPCC, a headline statement (high confidence) states that: “Under scenarios with increasing CO2 emissions, the ocean and land carbon sinks are projected to be less effective at slowing the accumulation of CO2 in the atmosphere” (IPCC, 2021: 36). Under the SSP1-1.9 (very low GHG emissions) scenario, the total cumulative CO₂ emissions taken up by land and ocean is projected to be 70% in 2100 (remaining 30% stays in the atmosphere). However, under the SSP5-8.5 (very high GHG emissions) scenario, this uptake reduces to 38% as ocean and land carbon sinks become less effective, meaning that there is likely to be a higher proportion of emitted CO2 remaining in the atmosphere (IPCC, 2021). In Chapter 5, AMS radiocarbon dates of these peatlands reveal that peatland initiation began at least ~ 7100 cal. yr BP; the implications are that these deposits may store a significant amount of carbon and have the potential to be an important part of the global carbon budget (Goyder et al., 2018). If these peatlands are disturbed or 109 degraded, carbon intake will be reduced, ultimately intensifying CO2 concentrations and further contribute to climate change, influencing the deposits in a negative, self- perpetuating cycle (Brevik and Homberg, 2004; Joosten 2009, 2011). In addition, if strong mitigation took place and GHG concentrations in the atmosphere were drastically reduced below current levels, historical emissions of long-lived GHGs will remain significant to future contributions of warming due to past accumulation and the inertia of the climate system (Skeie et al., 2021), and continue to impact these peatlands in future. Therefore, initial monitoring and preservation of the deposits are necessary in an uncertain climate future that depends on intact, natural carbon sinks (Friedlingstein et al., 2019). Peat is a soil distinguished from other soil types owing to the build-up of organic matter due to the combination of plant growth and waterlogging (Lindsay, 2010). Peat is therefore a direct product of the vegetation growing on the surface, reflecting prevailing hydrological and nutrient conditions controlled by climate and underlying landforms (Lindsay, 2010; Bain et al., 2011). In Chapter 6 and 7, valley vegetation EVI and NDVI and peatland NDVI indicated that vegetation growth has the same seasonality as the precipitation in the region. The implications are that vegetation occurring in the valley environment (which supports most of the peatland deposits classified in Chapter 5) is more strongly correlated with seasonal precipitation, and its growth is hydrologically dependent. Chapter 6 demonstrates that the Angolan Highlands vegetation was buffered against the drought periods that occurred since 2001. The NDVI and EVI timeseries indicates that the valley vegetation, which supports the peatlands, recovered at the start of each 110 rainfall season, a pattern which was identified even after the driest rainfall season on record during 2018/2019. In addition, although Chapter 7 did not include the drought data obtained from Chapter 6, the NDVI timeseries of the five intact peatland sites does overlap these drought periods. The peatland NDVI timeseries provide further evidence that the peatland vegetation is buffered against drought conditions, recovering at the start of each rainfall season even during drought periods. It is important that these peatlands remain undisturbed, as human activities would likely contribute to peatland degradation, limiting the peatland natural resistance to both drought and fire. In comparison to the valley vegetation region, miombo vegetation has peak greenness three months after peak precipitation. The ecological importance of the miombo woodlands cannot be overstated (Chiteculo and Surovy, 2018). The precipitation which falls on the upland environment, covered by Kalahari sand, is buffered by the miombo, and it is the interaction between the groundwater filtering through the sand and the adjacent seep line that allows and sustains peatland growth in this region, a mechanism for peat growth that has likely been present for millennia. In addition, the peatlands limit river flow and are a control valve between groundwater flow and the river as described in Chapter 5. The use of RS datasets provides contemporary environmental change indicators in a region which is historically war stricken and has had limited accessibility for scientific research (Conradie et al., 2016; Midgley and Engelbrecht, 2019). Chapters 6 and 7 demonstrate both direct and indirect anthropogenically induced environmental changes in the landscape. Over the 20-year fire record (2001-2020) presented in 111 Chapter 7, four common drought periods were identified within Chapter 6, of which three occurred since 2014. The two highest (2017 and 2019) annual burn area totals for the study area coincide with the 2017-2018 and 2018-2020 droughts, with 2019 having both the lowest seasonal rainfall total and highest burn area on record since 1981 and 2000, respectively. Increased burn area is likely due to the increased volume of dry material across the landscape, making fires more prevalent during periods of low rainfall and drought (Laris et al., 2016). Although fire occurrence could not be directly attributed to humans, the total burn area across the region highlights the potential influence of rural communities on fire, with an increase in burn area during 2003 following the end of the Angolan Civil War and a decrease during the 2020 lockdown associated with the COVID-19 pandemic. Precipitation deficits accounted for in the last 40 years indicate that rainfall has a negative trend, with the possibility of increased drought occurrence in future, consistent with what has been documented in Angola (Brooks et al., 2005; Cain, 2015; Carvalho et al., 2017) and throughout southern Africa (Abiodun et al., 2019; Nhamo et al., 2019; Gore et al., 2020). As a result of anthropogenic climate change, southern Africa, a region that is already characterised as dry and hot (Geppert et al., 2022), is projected to become generally drier under low-mitigation climate futures (Archer et al., 2018). 8.4 Concerns for the Angolan Highlands region Temperatures over southern Africa have increased rapidly over the last five decades, at a rate of near twice the global rate of temperature increase (Archer et al., 2018; 112 Gore et al., 2020). Under low-mitigation futures, further increases, up to 6°C by the end of the century relative to the present-day climate, may occur over the central and western interior regions (Engelbrecht et al., 2009; Archer et al., 2018; Abiodun et al., 2019). In projected changes of annual rainfall over southern Africa for the period 2080- 2099 compared to present day (1971-2000), the largest rainfall decreases are projected for Angola and the southern parts of South Africa (Archer et al., 2018). The projected decreases in Angola may be occu