Browsing by Author "Rubidge, Bruce"

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The affinities of the early cynodont reptile, Nanictosaurus
(Bernard Price Institute for Palaeontological Research, 1990) van Heerden, Jacques; Rubidge, Bruce
This investigation into the anatomy of the four extant specimens of Nanictosaurus has revealed that there is one valid species, viz. N. kitchingi Broom 1936 which has two junior synonyms, viz. N. robustus Broom 1940 and N. rubidgei Broom 1940. The closest known relative of Nanictosaurus is the well-known cynodont Thrinaxodon liorhinus. The differences from Thrinaxodon and other early cynodonts are discussed and illustrated.
A new GIS-based biozone map of the Beaufort Group (Karoo Supergroup), South Africa
(Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, 2010-12) van der Walt, Merrill; Day, Michael; Rubidge, Bruce; Cooper, Antony K.; Netterberg, Inge
The stratigraphic subdivision of the Karoo Supergroup of South Africa has been addressed for more than a century by various geoscience studies. A lack of good lithostratigraphic markers in the Beaufort Group and the fact that the succession preserves a rich diversity of therapsid fossils for which there is a robust taxonomic scheme, make biostratigraphy the best option for correlation. This is applicable both within the basin and when comparing the Karoo succession to other coeval depositional basins. Published biostratigraphic maps of the Beaufort Group were compiled based on rough estimates of the distribution of zone-defining tetrapod fossil genera throughout the basin. None of these simultaneously utilized all the databases of South African museum collections. The recent application of GIS technology to integrate the vertebrate fossil databases of all South African museums has resulted in a far more precise biozone map which can be continuously updated as new information is received. This digital map introduces an entirely new way of representing the geographical distribution of fossil data, and thus can potentially enhance basin development and tetrapod biogeographic studies.
Reassessment of the Phylogeny of Basal Therapsida Using Micro-Tomography and Bayesian Phylogeny
(University of the Witwatersrand, Johannesburg, 2023) Duhamel, Alienor; Rubidge, Bruce; Benoit, Julien
This PhD research is focussed on the evolution and cranial anatomy of the earliest therapsids. The Therapsida is a paraphyletic group of land tetrapods that lived mostly during the Permian and Triassic Periods. They emerged quite suddenly in the fossil record during the middle Permian, already showing diversification into five distinct groups: Biarmosuchia, Dinocephalia, Anomodontia, Gorgonopsia and Therocephalia. A sixth group, the Cynodontia, appeared during the late Permian. Of these, the anomodonts, therocephalians, and cynodonts survived the devastating Permo-Triassic mass extinction and continued to be the dominant tetrapods during the Early and Middle Triassic Periods. Ultimately, the cynodonts would evolve into the earliest mammals. Chapter 1 introduces the scientific questions tackled in this thesis. Chapter 2 provides the necessary explanations about the material and methods used for this work. The use of CT-scanning technology allows for the detailed examination of cranial internal morphology, with the goal of incorporating inner cranial characters into a phylogenetic framework. Chapters 3 and 6 focus on the fossilisation gap at the early to middle Permian transition, which has led to debates over the timing of the early diversification of the Therapsida. In Chapter 3, the geologic position and anatomy of one of the oldest known therapsids, Raranimus dashankouensis is reviewed. The chapter concludes with a reassessment of the basal phylogenetic position of Raranimus and suggests a Roadian origin for the taxon. Chapter 6 presents a comprehensive analysis of cranial morphological characters based on the descriptions and findings from Chapters 3, 4, and 5. Using both traditional maximum parsimony and clock-based Bayesian inference of phylogeny for the first time on the Therapsida, the results suggest that therapsids originated around 280.5 million years ago and rapidly diversified into several distinct clades during the Kungurian and Roadian Epochs. The phylogenetic relationships of several taxa are reevaluated and the findings suggest that Biseridens is more closely related to the Dinocephalia and Biarmosuchia, rather than the Anomodontia. Furthermore, the results suggest that Sinophoneus may represent the basal-most dinocephalian, and that Therocephalia might be paraphyletic. Therapsids were characterised by a wide range of cranial morphologies, with the Biarmosuchia and Anomod ontia exhibiting distinct cranial features. Both groups are considered basal among therapsids. Chapter 4 of this PhD research focuses on the postnatal ontogenetic development of cranial ornamentation in Biarmosuchia and possible impact on phylogeny. The results suggests that cranial bosses and ridges are ontogenetic features and the parietal bone originates from multiple centres of ossification. Chapter 5 is a comprehensive cranial description of several basal anomodont specimens. The Chapter concludes with a review of the taxonomy of the genus Eodicynodon and proposes the creation of a new taxonomical unit for specimen NMQR 2913.